The Cambridge natural history, Vol. 04 (of 10)Warburton, Cecil
Science
The Cambridge natural history, Vol. 04 (of 10)
Warburton, Cecil
Animals
The creature appears to be confined to the deep pools of the rivers and
tarns on the mountains of the southern and western portions of
Tasmania.[92] The waters in which it occurs are always cold and
absolutely clear, and there is no record of its living at altitudes much
below 2000 feet, while it frequently occurs at 4000 feet. The body may
attain upwards of two inches in length; it is deeply pigmented with
black chromatophores, and it is held perfectly horizontal without any
flexure. The animal rarely swims unless disturbed, usually walking about
on stones and water-plants at the bottom of deep pools. In walking the
endopodites of the thoracic limbs are chiefly instrumental, but they are
assisted by the exopodites of the abdominal limbs.
When frightened the shrimp can dart rapidly forwards or sideways by the
strokes of its powerful tail-fan, but it never jumps backwards as do the
other Malacostraca. It appears to browse upon the algal slime covering
the rocks and on the submerged liver-worts and mosses, but it does not
refuse animal food, even feeding on the dead bodies of members of its
own species. The thoracic limbs, which are all biramous except the last
pair, carry a double series of remarkable plate-like gills on their
coxopodites. The slender and setose exopodites of the thoracic limbs are
respiratory in function, being kept in continual motion even when the
animal is at rest, and serving to keep up a current of fresh water round
the gills.
_Anaspides_ shows a remarkable combination of structural characters,
some of which are peculiar, while others are possessed in common with
the Peracarida or Eucarida. The chief peculiar characters are the entire
absence of a carapace, and the freedom of the eight thoracic segments,
with eight free thoracic ganglia in the nerve-cord; the peculiar double
series of plate-like gills; the structure of the alimentary canal; and
the fact that the eggs, instead of being carried in a brood-pouch, or
affixed to the abdominal limbs, are deposited under stones and among
water-plants.[93]
[Illustration:
FIG. 77.—_Anaspides tasmaniae_ in natural position for walking, × 1.
The last two pereiopods point backwards and are overlapped by the
first two pleopods.
]
The Peracaridan features, uniting it especially with the Mysidacea, are
the structure of the elongated heart, the filiform spermatozoa, and the
fact that no complicated metamorphosis is passed through, the young
hatching out in a condition similar to, though possibly not identical
with, the adult form.
The Eucaridan, especially Decapodan, features are the presence of an
auditory sac on the basal joint of the antennules, and the modification
of the endopodites of the first two abdominal appendages in the male to
form a copulatory organ.
A type of a new genus of this family was found by me in the littoral
zone of the Great Lake of Tasmania at an elevation of 3700 feet, and
named _Paranaspides lacustris_.
Public-domain text, read in full here on John Shaqi.
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