The Cambridge natural history, Vol. 04 (of 10)Warburton, Cecil
Science
The Cambridge natural history, Vol. 04 (of 10)
Warburton, Cecil
Animals
_Sao hirsuta_ is a species found in the Cambrian, the development of
which was fully described by Barrande. Its earliest protaspis stage
(Fig. 143, A) is circular in outline; the glabella expands in front and
reaches the anterior margin; the pygidial region is not distinctly
separated from the cephalic region; segmentation is indicated in the
former, and the neck-ring is present in the latter; the eye-line is seen
on each side of the glabella near the anterior margin. In a later stage
(Fig. 143, C) the segmentation of the glabella becomes more distinct,
indicating the existence of five cephalic segments, and the facial
suture appears near the margin limiting a very narrow free cheek.
Subsequently (Fig. 143, D-F) the thoracic segments develop, and increase
in number until the adult stage (G) is reached; also the eyes appear at
the margin of the cephalic shield, and gradually move inwards, and the
glabella becomes narrower and rounded in front, and ceases to reach the
anterior margin. In this species the eye-line is present in the adult.
[Illustration:
FIG. 144.—_Triarthrus becki_, Green. Ordovician. =A=, =B=, Two
successive stages of the protaspis, × 45. (After Beecher.)
]
In the protaspis of _Triarthrus_ (Fig. 144), found in the Ordovician,
the glabella does not reach the front margin nor expand in front as it
does in _Sao_; an eye-line is present, but disappears before the adult
stage is reached.
[Illustration:
FIG. 145.—Larval stages of Trilobites. =A-D=, _Dalmanites socialis_,
Barr. Ordovician, Bohemia. The small figures below show the natural
size of each specimen. (After Barrande.) =E=, _Mesonacis
asaphoides_, Emmons, × 10. Lower Cambrian, North America. (After
Walcott.) =F=, _Acidaspis tuberculata_, Conrad, × 20. Lower
Helderberg Group (Lower Devonian or Upper Silurian), Albany County.
(After Beecher.)
]
_Dalmanites_ (Fig. 151, C) is a more advanced type than _Sao_ and
_Triarthrus_, and is found in later deposits. In the earliest stage
(Fig. 145, A) the head and pygidium are quite distinct, and there is no
eye-line present at this or any stage in development, but large ovoid
eyes are found on the front margin, and have their long axes placed
transversely to the axis of the body; the glabella is strongly segmented
and is rounded in front. In later stages (C, D) the pygidium increases
in size relatively, and the thoracic segments are successively
introduced; the facial sutures and free cheeks appear on the dorsal
surface, and as the free cheeks grow the eyes move inwards and
backwards, and gradually swing round until their long axes become
parallel with the axis of the body.
The larval form of _Acidaspis_ (Fig. 145, F) is of interest since even
in the earliest stage it shows the spiny character which forms such a
striking feature of the adult (Fig. 151, F).
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