The Cambridge natural history, Vol. 04 (of 10)Warburton, Cecil
Science
The Cambridge natural history, Vol. 04 (of 10)
Warburton, Cecil
Animals
_Limulus_ is an example of a persistent type. It appears first in
deposits of Triassic age, and is found again in the Jurassic,
Cretaceous, and Oligocene. In the lithographic limestone of Solenhofen
in Bavaria, which is of Upper Jurassic age, _Limulus_ is common and is
represented by several species. One species is known from the Chalk of
Lebanon, and another occurs in the Oligocene of Saxony. No other genus
of the Xiphosura appears to be represented in the Mesozoic and Tertiary
deposits, but in the Palaeozoic formations (principally in the Upper
Silurian, the Old Red Sandstone, and the Coal Measures) several genera
have been found, most of which differ from _Limulus_ in having some or
all of the segments of the abdomen free; in this respect they resemble
the Eurypterida, but differ from them in the number of segments. In
_Hemiaspis_ (Fig. 159, A), from the Silurian, the segments of the
abdomen are divisible into two groups (mesosoma and metasoma) in the
same way that they are in Eurypterids; the first six segments have
broad, short terga, the lateral margins of the sixth being divided into
two lobes, probably indicating the presence of two fused segments; the
last three segments are narrower and longer than the preceding, and at
the end is a pointed tail-spine. In _Belinurus_ (Fig. 160) from the
Carboniferous, the two regions of the abdomen are much less distinct;
there are eight segments, the last three of which are fused together,
and a long tail-spine. In _Neolimulus_, from the Silurian, there seems
to be no division of the abdomen into two regions, and apparently all
the segments were free. On the other hand, in _Prestwichia_
(Carboniferous), all the segments of the abdomen, of which there appear
to be seven only, were fused together (Fig. 159, B).
[Illustration:
FIG. 160.—_Belinurus reginae_, Baily, Coal Measures, Queen’s Co.,
Ireland, × 1. (After Woodward).
]
In the Palaeozoic genera the median or axial part of the dorsal surface
is raised and distinctly limited on each side, so presenting a trilobed
appearance similar to that of Trilobites. In _Neolimulus_, _Belinurus_,
and _Prestwichia_, lateral eyes are present on the sides of the axial
parts of the carapace, and near its front margin median eyes have been
found in the two last-named genera.
In nearly all the specimens of Palaeozoic Xiphosura[225] which have been
found nothing is seen but the dorsal surface of the body; in only a very
few cases have any traces of the appendages been seen,[226] but, so far
as known, they appear to have the same general character as in
_Limulus_.
_Aglaspis_, found in the Upper Cambrian of Wisconsin, has been regarded
as a Xiphosuran. If that view of its position is correct, then
_Aglaspis_ will be the earliest representative of the group at present
known. Other genera of Palaeozoic Xiphosura are _Bunodes_, _Bunodella_,
and _Pseudoniscus_ in the Silurian; _Protolimulus_ in the Upper
Devonian; and _Prolimulus_ in the Permian.
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