The Cambridge natural history, Vol. 06 (of 10)Sharp, David
Science
The Cambridge natural history, Vol. 06 (of 10)
Sharp, David
Animals
This enormous series includes about 25,000 species, and as may well be
imagined shows a great variety of structure amongst its forms. The vast
majority may, however, be readily recognised by the two characters
mentioned above. There are some cases in which the beak is indistinct, and
others in which the tarsi are {289}five-jointed (_Dryophthorus_), and even
slender (Platypides). In these cases a close examination shows that the
gular region on the middle of the back of the under surface of the head
cannot be detected, and that the back of the prosternum is very strongly
consolidated by the side-pieces of the thorax meeting together and being
very firmly joined behind the coxae. The beak is in the great majority
perfectly distinct, though it varies so extremely in form that it can only
be briefly described by saying that it is a prolongation of the head in
front of the eyes, or that the antennae are inserted on its sides near to,
or far from, the tip. It has been ascertained in many cases that the
rostrum is used by the female to assist in placing the eggs in suitable
places, a hole being bored with it; in some cases it is also used to push
the egg far into the hole in which it has previously been placed by the
ovipositor; but there are many forms in which it is fairly certain that it
is not so used. What purpose it serves in the male is totally unknown. In
many members of the series, the rostrum differs in form in the two sexes,
and in most, if not in all, these cases it is clear that the distinctions
tend in the direction of making the beak of the female more efficient for
the mechanical purpose we have mentioned.
[Illustration: Fig. 147.—_Eugnoristus monachus_ ♀. Madagascar. A, The
imago; B, front of pronotum, head, and rostrum.]
It was proposed by Leconte and Horn to separate this series from all the
other Coleoptera as a primary division, and they looked on it as of lower
or more imperfect structure. Packard has very properly protested against
this interpretation; and there seems to be no reason whatever for
considering the Rhynchophora as "lower" than other beetles; indeed we
should be inclined to place such forms as Calandrides {290}amongst the most
perfect of Insects; their external structure (as shown by _Eugnoristus
monachus_, Fig. 147) being truly admirable.
Only four families of Rhynchophora can be at present accepted as
satisfactory; one of these—Curculionidae—includes an enormous majority of
the whole series. Though it is probable that it will ultimately be divided
into several families, the attempts to that end that have already been made
are not satisfactory.
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