The Cambridge natural history, Vol. 08 (of 10)Gadow, Hans
Science
The Cambridge natural history, Vol. 08 (of 10)
Gadow, Hans
Animals
Considering the age of these little creatures and their low position in the
reptilian scale–in fact, they stand almost as low {289}as the
Proreptilia–it is not to be wondered at that they still retain a number of
amphibian features. The skull is encased in dermal bones as in the
Stegocephali, and the dermal armour of the trunk and tail is composed of
many bony, sculptured scales, which cover back, sides, and under surface.
The middle rows on the back are the largest, while the scales on the belly
are arranged in transverse rows, which imbricate and converge obliquely
headwards. Special gular plates seem to be absent. The skull has an
interparietal foramen. The jaws and the palate are furnished with small,
simple teeth, and there is a large parasphenoid bone, an eminently
amphibian character. The occipital condylar articulation is supposed to be
double. The centra of the vertebrae are deeply amphicoelous, elongated, and
constricted in the middle, just like those of the Aistopoda and
Branchiosauri. The dorsal spinous processes are strongly developed, and
with the zygapophyses are very reptilian. Transverse processes are absent
or very short, the tubercular portions of the ribs articulating with the
centra, the capitula mostly intervertebrally, in any case close to the
anterior end of the centra. The tail-vertebrae possess very typical,
movable chevrons, placed intervertebrally, and bear an extraordinary
resemblance to those of Geckos. The ribs are long and slender, but there is
no sternum. The fore- and hind-limbs are pentadactyle, in opposition to the
invariably four-fingered Stegocephali. The shoulder-girdle consists of
scapulae, coracoids, clavicles, cleithra, and a T-shaped interclavicle. The
pelvis also resembles that of certain Stegocephali by the separately
ossified, somewhat disc-shaped, flat ischia and pubes, which seem to have
been joined together by cartilage into one broad mass.
_Hylonomus_, Dawson's type of Microsauri, was found in the Coal-measures of
Nova Scotia, within decayed tree-stumps. Closely allied, if not identical,
but much better known is _Hyloplesion_, e.g. _H. longicostatum_ of the
uppermost Permian of Nyrschan in Bohemia. Total length under 4 inches; eyes
with bony sclerotic rings; neck short. The truly Permian genera _Dawsonia_,
_Melanerpeton_, _Orthocosta_, and _Seeleya_ are allied forms, the last
scarcely one inch in length, but well preserved. _Petrobates_ of the
Triassic Lower Red Sandstone of Saxony has an arrangement of the ventral
dermal armour closely resembling abdominal ribs.
{290}ORDER II. PROSAURI
Mostly extinct, chiefly Permian and Triassic, terrestrial, unarmoured
reptiles with deeply biconcave vertebrae, numerous intercentra and
chevron-bones, fixed quadrates, complete pentadactyle limbs and
shoulder-girdle, entepicondylar foramina, acrodont teeth, and many small
abdominal ossifications.
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