The Cambridge natural history, Vol. 08 (of 10)Gadow, Hans
Science
The Cambridge natural history, Vol. 08 (of 10)
Gadow, Hans
Animals
Probably several other genera of American Permian and also of European
Permian strata will, when better known and critically examined, have to be
referred to the Proreptilia. Thus for instance the European _Melosaurus_
may have affinities with _Eryops_, while _Diplovertebron_ of Bohemia seems
to be allied to _Cricotus_. The difficulty of division will lie with those
Lower Permian Amphibia which, like _Archegosaurus_, _Euchirosaurus_,
_Actinodon_, possess tripartite vertebrae, which at first sight are
strikingly like those of _Eryops_. But the tail-vertebrae permit of no
mistake, and since these are quadripartite in _Archegosaurus_,
_Chelydosaurus_, and _Sphenosaurus_, these genera are safely to be classed
with the Amphibia, unless, indeed, for mere argument's {288}sake, it be
assumed that the intercentral discs of _Diplovertebron_ and _Cricotus_ are
formed by the fusion of Amphibian interdorsals with interventrals. Anyhow,
simply to state that the tripartite vertebrae of _Eryops_ are the same as
those of _Actinodon_, would be as convincing as saying that the English and
French flags are essentially the same, both containing the same colours,
but one is white, red, and blue, the other blue, white, and red. Tripartite
Amphibian vertebrae are composed of basidorsals + basiventrals +
interdorsals, those of Reptiles are made up of basidorsals + basiventrals +
interventrals. (Cf. Fig. 56, p. 283, and Fig. 1, p. 13.)
_SUB-CLASS II.–PROSAURIA._
_Mostly extinct Reptiles, with deeply amphicoelous but stereospondylous
vertebrae, with movable chevron-bones in the tail and frequently with
intercentra in the trunk._ Sphenodon, _the only recent genus, has no
copulatory organs_.
ORDER I. MICROSAURI.
_Extinct, small Reptiles, mostly Carboniferous and Permian, with dermal
armour on the dorsal and ventral side and with bifurcated ribs._
We retain this term of Dawson's for those small, newt-shaped, chiefly
Permian reptiles, which are allied to _Hylonomus_, after elimination of
contemporary forms like _Keraterpeton_ and _Urocordylus_, which belong to
the Branchiosaurian order of the Stegocephali. Until recently[122] all
these creatures had been classed with the Stegocephali. The Microsauri in
the present restricted sense reveal themselves, however, as reptiles by the
movable chevron-bones in their tail, their broad neurocentral sutures, the
possession of two sacral vertebrae (_Petrobates_), the bifurcated ribs
which always articulate with the centra (most clearly shown in
_Orthocosta_), and the possession of five fingers and toes.
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