The Cambridge natural history, Vol. 08 (of 10)Gadow, Hans
Science
The Cambridge natural history, Vol. 08 (of 10)
Gadow, Hans
Animals
Total length of very large male specimens up to two feet and a half; in
general appearance like many a stoutly built lizard. The general colour of
the skin is dark olive-green with small white or yellowish specks on the
sides. A series of slightly erectile spines of yellowish colour extends
from the top of the head to the end of the tail, but is interrupted on the
neck; they are cutaneous, covered with a thin sheath of horn. The under
surface is covered with numerous scales, arranged in transverse rows; the
rest of the body is rather granular. The tail is thick, slightly compressed
laterally. The eye is large, dark brown, with a vertical pupil.
Those who are satisfied with superficial resemblances still group this
creature with the lizards, but it reveals itself as a primitive reptile or
Prosaurian by the following characters, every one of which distinguishes it
from the lizards:–The temporal region is bridged by three bony arcades. The
large vomers, palatines, and pterygoids form a broad bony roof to the
mouth; the large quadrates are firmly fixed by the pterygoids, squamosals,
lateral occipital bones, and by the jugal bridge. The vertebrae possess an
unbroken series of intercentral wedge-bones. There is an elaborate system
of abdominal ribs. The humerus has an entepicondylar foramen, and there is
also, in contradistinction to the fossil Rhynchocephalia, an ectepicondylar
foramen for the passage of the radial nerve. The carpus still has the
primitive number of ten bones, all of which remain separate, including the
intermedium. Of soft parts are to be mentioned above all the entire absence
of external copulatory organs, _Sphenodon_ being the {295}only recent
reptile which is devoid of them; a most primitive condition, sufficient by
itself to separate this creature from all the other living reptiles.
[Illustration: FIG. 59.–A, Dorsal; B, ventral; C, left-sided view of the
skull of _Sphenodon_. × 3/2. _Col_, columella auris; _Cond_, occipital
condyle; _E.P_, ectopterygoid; _F_, frontal; _Jug_, jugal; _Max_,
maxillary; _Na_, nasal; _No_, anterior nasal opening; _Pal_, palatine;
_Par_, parietal; _Pmx_, premaxillary; _Prf_, prefrontal; _Pt.f_,
postfrontal and postorbital; _Ptg_, pterygoid or endopterygoid; _Q_,
quadrate and quadrato-jugal; _Sq_, squamosal; _Vo_, vomer. See also Fig.
54, B, p. 280.]
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