The Cambridge natural history, Vol. 08 (of 10)Gadow, Hans
Science
The Cambridge natural history, Vol. 08 (of 10)
Gadow, Hans
Animals
The dentition is most abnormal, and permits the division of the Theromorpha
into two or three main groups. In the Pareiasauri the teeth of the upper
and lower jaws form rather even series of nearly equal size; smaller teeth
are carried by the palatal bones. In the Theriodontia the teeth are
differentiated in a truly Mammalian fashion into incisors, prominent
canines, and multicuspid or tubercular molars. Each tooth, and this applies
to all Theromorpha, is implanted in a separate alveolus; _Tritylodon_ only
seems to have double-rooted molars. The lower canines cross in front of the
upper, just as in Mammals. In _Placodus_, which probably belongs to this
assembly, the teeth are few in numbers, very broad and flat, especially
those of the palate. In _Dicynodon_ and _Gordonia_ the teeth are restricted
to a pair of conical, sometimes very large, tusk-like upper canines, and in
_Oudenodon_ the whole mouth is toothless.
The configuration of the skull shows two main types. In the Pareiasauri it
is completely roofed in by dermal bones, the only holes on the surface
being the nostrils, orbits, and the interparietal foramen.
The most striking feature of the second type of skull is the tendency to
form an almost Mammalian zygomatic arch by the junction of the much
elongated squamosal with the jugal bone, both abutting against a downward
process of the postfrontal bone. The skull shows a pair of wide
supratemporal foramina bordered by the parietals, squamosals, and
postfrontals. The composition of the temporal arch varies considerably in
detail, and in _Cynognathus crateronotus_ at least there is a small hole
within the arch, between the squamosal and jugal, probably the last remnant
of the otherwise absent infratemporal foramen. Except in the roofed-in
skulls of _Pareiasaurus_ and _Elginia_ there is no separate quadrato-jugal
element. The quadrate is firmly fixed by the overlapping squamosal, and the
whole pedicle for the support of the mandible is rather elongated, and
either stands vertically or slants forwards. The mandible itself is
compound. The pterygoids extend backwards so as to approach or reach the
{302}distal portion of the quadrate; separate ectopterygoids do not seem to
be developed. The shoulder-girdle consists on either side of a large
scapula, which is mostly directed obliquely backwards, and is fused with
the coracoid; a precoracoid is present or at least indicated by a notch or
foramen; it is usually fused with the other bones. At least some genera
possess a T-shaped interclavicle and clavicles; _Pareiasaurus_ possesses
also a pair of cleithra.
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