The Cambridge natural history, Vol. 08 (of 10)Gadow, Hans
Science
The Cambridge natural history, Vol. 08 (of 10)
Gadow, Hans
Animals
Teeth are more restricted in their occurrence than in the Urodela. On the
jaws they always stand in one row. With the exception of the
Hemiphractinae, Amphignathodontinae, Ceratobatrachinae, and Genyophryninae,
no recent Anura carry teeth on the lower jaw, and even in these genera they
are mostly much reduced in size and firmness, having all the appearance of
vanishing structures. The premaxillae and maxillae are frequently furnished
with teeth, except in the Dendrobatinae, Genyophryninae, Engystomatinae,
Dendrophryniscinae, Bufonidae, _Pipa_, and _Hymenochirus_. The vomers
mostly carry a series of teeth on their posterior border; when these teeth
are absent, as in many species of _Bufo_, a kind of substitute sometimes
occurs on the palatines in the shape of a row of tuberosities. The
palatines carry teeth in Hemiphractinae. The parasphenoids are rarely
toothed, _e.g._ _Triprion_, _Diaglena_, _Amphodus_, and occasionally in
_Pelobates_.
A few Anura possess peculiar substitutes for teeth in the anterior portion
of the lower jaw, namely, a pair of conical bony processes, sometimes
rather long, but always covered by the dense gums, or investment of the
jaws; _e.g._ _Lepidobatrachus_, several Rana, _e.g._ _R. adspersa_, _R.
khasiana_, _R. kuhli_, and _Cryptotis brevis_.
{31}Cranial dermal ossifications are developed in some species of _Bufo_,
still more in the Hemiphractinae, and above all in _Pelobates cultripes_
and in the Cystignathoid genus _Calyptocephalus_.
The HYOID APPARATUS of the Anura is complicated. It is originally composed
of the hyoidean and four branchial arches, with one median, copular piece.
The branchial arches form in the early life of the tadpole the elaborate
framework of the filtering apparatus mentioned on p. 44. During
metamorphosis the whole filter disappears, owing to resorption of the
greater part of the branchial arches; only their median portions remain,
and fuse with the enlarged copular piece and the hyoidean arches into a
broad shield-shaped cartilage (_corpus linguae_), whence several lateral
processes sprout out, the posterior pair of which are generally called
thyrohyals or thyroid horns. The true hyoid horns give up their larval
lean-to articulation with the quadrate, become greatly elongated, and gain
a new attachment on the otic region of the cranium. The transformation of
the whole apparatus has been studied minutely by Ridewood, in _Pelodytes
punctatus_.[13]
SKIN
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