The Cambridge natural history, Vol. 09 (of 10)Evans, A. H. (Arthur Humble)
Science
The Cambridge natural history, Vol. 09 (of 10)
Evans, A. H. (Arthur Humble)
Animals; Birds
9. The organs of deglutition and digestion begin with the tongue, which is
subject to much variation of structure, according to the different groups
of Birds, and is of course correlative with their habits. It has little
connexion with taste, though often of assistance in obtaining nutriment. To
this follows the gullet (_oesophagus_), which in many cases has an
enlargement forming the crop (_ingluvies_), wherein the food may be
temporarily retained before passing into the stomach, the last-named always
having an antechamber (_proventriculus_) where digestion is largely
accomplished, in front of the gizzard (_ventriculus_). This has frequently
strong muscular walls, and its action is often assisted by the mechanical
process of comminution performed by stones, grit or sand, swallowed for
that purpose. The stomach is succeeded by the intestines, which in most
cases have a pair of blind-sacs (_caeca_) attached to them, often acting as
aids to digestion, though these are not always functional, and are absent
in many Birds, while in others they attain a very large size, their
condition being in consequence of some importance as a systematic
character.
10. The organs of voice in Birds have long attracted special interest from
the loud cries which some utter, and the melody with which others are
gifted. Setting aside the part played by {13}the _trachea_ or windpipe in
supplying air to the lungs, its formation is worthy of attention. Its upper
end consists of the _larynx_, and it passes down the neck as a flexible
tube, formed by a continuous succession of bony rings connected by
membrane, until it bifurcates into two _bronchi_, which open into the
lungs. A common feature, found in many groups not nearly allied, is the
dilatation of a portion, generally near the middle, while a remarkable
modification is exhibited by the males of many of the Duck-tribe, some of
the lowest rings being fused together and forming what is known as the
_bulla ossea_ or "labyrinth." In other _Anatidae_ (some of the Swans), and
some of the Cranes, the trachea enters the keel of the sternum; but a not
unfrequent modification, usually confined to the male sex, often occurs
elsewhere, when the windpipe is looped back upon itself. All these
arrangements, however they may affect the sounds uttered by Birds, do not
in themselves constitute the voice organ of most. That is reserved for the
_syrinx_, a peculiarity of the Class _Aves_, consisting of the lower end of
the trachea and the adjoining part of the bronchial tubes; and the varied
modulations are effected by means of muscles attached thereto. These
voice-muscles may be wholly absent or of the simplest character, but they
attain their highest perfection in the _Passeres_, and especially in the
large group of them known as _Oscines_, where there are often five or seven
pairs. In this group the lowest four or five tracheal rings are solidly
fused into a little bony box communicating with the bronchi; the first and
Public-domain text, read in full here on John Shaqi.
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