The Cambridge natural history, Vol. 09 (of 10)Evans, A. H. (Arthur Humble)
Science
The Cambridge natural history, Vol. 09 (of 10)
Evans, A. H. (Arthur Humble)
Animals; Birds
This genus includes the species _P. longissimus_, _P. inflatus_, _P.
platygnathus_, _P. modicus_, _P. gracilis_, and _P. sehuensis_;
_Brontornis_, which has a shorter and wider mandible and smaller but
stouter metatarsi, possesses in _B. burmeisteri_ a form as large as
_Aepyornis maximus_, while _Opisthodactylus_ and other proposed genera are
too imperfectly known to deserve consideration in our limited space.
Besides the above, Dr. Gadow classed with the Stereornithes, _Diatryma_ of
New Mexico, known from a metatarsus; _Dasornis_ of the London Clay,
described from fragments of a skull; _Remiornis_ from the neighbourhood of
Rheims, of which several imperfect bones have been found; and _Gastornis_
of both England and France, of which a fair number of parts have been
unearthed. All occur in the Eocene, but the question of their relationship
is by no means settled, and some writers consider _Gastornis_ to be nearly
allied to the Anseres. This form appears to have been of the size of an
Ostrich, with long leg-bones and short weak wings, and was probably
flightless. Three species have been propounded, _G. parisiensis_, _G.
klaasseni_, and _G. edwardsi_.
* * * * *
(B) With regard to the difficult question of the position in the system of
the NEORNITHES ODONTOLCAE, a few introductory words of explanation are
necessary. In 1872 Professor Marsh bestowed upon two fossils from the
Cretaceous deposits of Kansas the names of _Hesperornis_ and _Ichthyornis_,
which he proposed in the following year[45] to comprise in a Sub-class
_Odontornithes_, so called from the presence of teeth in the jaws.
Subsequently[46] he divided this Sub-class into two Orders, _Odontolcae_
and _Odontotormae_, the former containing _Hesperornis_, with the teeth
arranged in grooves, and the latter _Ichthyornis_, where they were placed
in distinct sockets. His views have been controverted by many writers, but
Mr. Lydekker–an authority of great weight in this connexion–while fully
admitting the affinity of the first form to {46}the Divers, and the
resemblance of the second to the Gull-tribe, proposed in 1891[47] to retain
the term Odontornithes for a series of birds ancestral to the modern series
of toothless Carinatae, for which he adopted the title _Euornithes_, used
in a narrower sense by Dr. Stejneger. It has, however, been decided to
follow Dr. Gadow on this point; while the marks of distinction given below
make it seem at least probable that, whereas _Ichthyornis_ may be referred
to the Carinate division, _Hesperornis_ should be placed in closer
proximity to the Ratite forms. Our Neornithes Odontolcae consequently
contain the HESPERORNITHES, the ENALIORNITHES, and _Baptornis_, all of
which appear to be nearly related.
[Illustration: FIG. 13.–Restoration of _Hesperornis_. (From Huxley, after
Marsh.) × 1/13.]
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