The Cambridge natural history, Vol. 10 (of 10)Beddard, Frank E. (Frank Evers)
Science
The Cambridge natural history, Vol. 10 (of 10)
Beddard, Frank E. (Frank Evers)
Animals; Mammals
Scent is no doubt a sense of higher importance in mammals
than in birds. The subject is one which will bear further study.
NAILS AND CLAWS.--Except for the Cetacea (where rudiments have been found
in the foetus), the extremities of the fingers and of the toes of mammals
are covered by, or encased in, horny epidermic plates, known as nails,
claws, and hoofs.
The variety in the shape and development of these corneous sheaths to the
digits is highly characteristic of mammals as opposed to lower Vertebrates.
If we take extreme cases, such as the nail of the thumb in Man, the hoof of
a Horse, and the claw of a Cat, it is easy to distinguish the three kinds
of phalangeal horny coverings. But the differences become extinguished as
we pass from these to related types. The nail of the little finger in Man
approaches the claw-like form; and the hoofs of the Lama are almost claws
in the sharpness of their extremities. On the whole it may be said that
claws and hoofs embrace the bone which they cover, while nails lie only
upon its dorsal surface. The form of the distal phalanx which bears the
nail shows, however, two kinds of modification which do not support such a
classification. When those phalanges are clad with hoofs or covered by a
nail they end in a rounded and flattened termination. On the other hand,
when they bear a claw they are themselves sharpened at the extremity and
often grooved above.
THE MARSUPIUM.--It may appear to be unnecessary at this juncture to speak
of the marsupial pouch, which is so usually believed to be a characteristic
of the group Marsupialia. Rudiments of this structure have, however, been
recently discovered in the higher mammals, and, as Dr. Klaatsch[6] has
remarked, all researches into the "history of the mammals culminate in the
question whether the placental mammals pass through a marsupial stage or
not." We cannot, therefore, look upon the marsupial pouch as a matter
affecting only the Marsupials, though it is true that this organ is at
present functional only in them and in the Monotremata. {15}
[Illustration] FIG. 3.--_Echidna hystrix._ A, Lower surface of brooding
female; B; dissection showing a dorsal view of the pouch and mammary
glands; ++, the two tufts of hair in the lateral folds of the mammary pouch
from which the secretion flows, _b.m_, Pouch; _cl_, cloaca; _g.m_, groups
of mammary glands. (From Wiedersheim's _Comparative Anatomy_, after W.
Haacke.)
In the Marsupials the pouch shelters the young, which are born in an
exceedingly imperfect state, minute, nude, and blind, with a "larval" mouth
fitted only to grasp in a permanent fashion the teat, upon which they are
carefully fixed by the parent. But even later the pouch is made use of as a
temporary harbour of refuge: from the pouch of female Kangaroos at the
Zoological Gardens may frequently be observed to protrude the tail and
hind-legs of a young Kangaroo as big as a Cat, and perfectly well able to
take care of itself.
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