Be that as it may, the evolutionist interprets the resemblances of
homology as surviving vestiges of an ancient ancestral type, which
have managed to persist in the descendants notwithstanding the
transformations wrought in the latter by the process of progressive
divergence. Moreover, just as the existence of a common ancestor is
inferred from the _fact_ of resemblance, so the relative position
in time of the common ancestor is inferred from the _degree_ of
resemblance. The common ancestor of forms closely allied is assumed to
have been proximate, that of forms but distantly resembling each other
is thought to have been remote. Thus the common ancestor of species
grouped under the same genus is supposed to have been less remote
than the common ancestor of all the genera grouped under one family.
The same reasoning is applied, _mutatis mutandis_, to the ancestry of
families, orders and classes.
The logic of such inferences may be questioned, but there is no
blinking the fact that, in practice, the genetic explanation of
homology is assumed by scientists to be the only reasonable one
possible. In fact, so strong is their confidence in the necessity of
admitting a solution of this kind, that they do not hesitate to make it
part and parcel of the definition of homology itself. For instance, on
page 130 of Woodruff’s “Foundations of Biology” (1922), we are informed
that homology signifies “a fundamental similarity of structure based on
descent from a common antecedent form.” The Yale professor, however,
has been outdone in this respect by Professor Calkins of Columbia, who
discards the anatomical definition altogether and substitutes, in lieu
thereof, its evolutionary interpretation. “When organs have the same
ancestry,” he says, “that is, when they come from some common part of
an ancestral type, they are said to be homologous.” (“Biology,” p.
165.) In short, F. A. Bather is using a consecrated formula culled from
the modern biological creed when he says: “The old form of diagnosis
was _per genus et differentiam_. The new form is _per proavum et
modificationem_.” (_Science_, Sept. 17, 1920, p. 259.)
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