Finally, it may be noted in passing that, though it is customary
with evolutionists to regard homologous characters as the tenaciously
persistent heritage of primeval days, and to look upon adaptational
characters as adventitious and accessory to the aforesaid primitive
heritage, the supposedly older and more fundamental characters fail to
give, by the manifestation of greater fixity, any empirical evidence
whatever of their being more deeply or firmly rooted in the hereditary
process than the presumably newer adaptational characters. We have,
therefore, no experimental warrant for appropriating homologous,
rather than adaptational, characters to the process of inheritance.
“It is sometimes asserted,” says Goodrich, “that old-established
characters are inherited, and that newly begotten ones are not, or are
less constant, in their reappearance. This statement will not bear
critical examination. For, on the one hand, it has been conclusively
shown by experimental breeding that the newest characters may be
inherited as constantly as the most ancient.... While, on the other
hand, few characters in plants can be older than the green color
due to chlorophyll, yet it is sufficient to cut off the light from
a germinating seed for the greenness to fail to appear. Again, ever
since Devonian times vertebrates have inherited paired eyes; yet,
as Professor Stockard has shown, if a little magnesium chloride is
added to the sea water in which the eggs of the fish _Fundulus_ are
developing, they will give rise to embryos with one median cyclopean
eye! Nor is the suggestion any happier that the, so to speak, more
deep-seated and fundamental characters are more constantly inherited
than the trivial or superficial. A glance at the organisms around
us, or the slightest experimental trial, soon convinces us that the
apparently least important character may reappear as constantly as
the most fundamental. But while an organism may live without some
trivial character, it can rarely do so when a fundamental character is
absent, hence such incomplete individuals are seldom met in Nature.”
(_Science_, Dec. 2, 1921, p. 530.)
But, whether it be upon, or beneath, the surface, similitude of _any
kind_ suffices to establish our contention that inheritance is not the
only similifying influence present in organisms, and that resemblance
is perfectly compatible with independence of ancestry. We have,
therefore, an alternative for inheritance in the explanation of organic
uniformities, and by the admission of this alternative, which, for the
rest, is factually attested by the universally acknowledged phenomena
of convergence, the inference of common descent from structural
resemblance is shorn of the last remnant of its demonstrative force, as
an _a posteriori_ argument.
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