The Chain of Life in Geological Time: A Sketch of the Origin and Succession of Animals and PlantsDawson, John William, Sir
Science
The Chain of Life in Geological Time: A Sketch of the Origin and Succession of Animals and Plants
Dawson, John William, Sir
Paleobotany; Paleontology
It is curious that in the line of succession above stated, the beautiful
tubulated cell-wall of _Eozoon_ disappears; and this structure seems,
after the Laurentian, to be for ever divorced from the great laminated
Protozoans. It reappears in the Carboniferous, in certain smaller
organisms of the type of the _Nummulites_, or Money-stone Foraminifers,
and is continued in this group of smaller and free animals down to the
present time. In the Cretaceous and early Tertiary periods, the
Foraminifera of different types have been nearly as great rock-builders
as they were in the Laurentian. Some of these later rock-builders,
however, have belonged to the lower or imperforate group; others to the
higher or Rotaline and Nummuline groups; and, as a whole, they have been
individually small, making up in numbers what they lacked in size.
Probably the conditions for enabling animals of this type rapidly, and
on a large scale, to collect calcareous matter, were more favourable in
the Laurentian than they have ever been since.
[Illustration: FIG. 24.—_Archæocyathus minganensis._ A Primordial
Protozoon.—After Billings.
_a_, Pores of the inner wall.]
In the Siluro-Cambrian age two other forms of gigantic Foraminiferal
Protozoans were introduced, widely different from _Eozoon_, and destined
apparently not to survive the period in which they appeared. These were
_Archæocyathus_, the ancient Cup-corals, and Receptaculites, which may
perhaps be called the Sack-corals. Both are quite remote from _Eozoon_ in
structure, wanting its complexity in the matter of minute tubules, and
having greater regularity and complication on the large scale.
_Archæocyathus_ had the form of a hollow inverted cone with double
perforated walls, connected by radiating irregular plates, also
perforated (Fig. 24). It has been regarded as a sponge, and some species
are certainly accompanied with spicules; but these I have ascertained to
be merely accidental, and will be referred to in the next chapter. The
true structure of _Archæocyathus_ consists of radiating calcareous plates
enclosing chambers connected by pores. Archæocyathus came in with the
Later Cambrian, and seems to have died out in the Siluro-Cambrian. The
only more modern things which at all resemble it are the Foraminifera
called _Dactylopora_, which belong to the Tertiary period.
[Illustration: FIG. 25.—Receptaculites. Restored.—After Billings.
_a_, Aperture.
_b_, Inner wall.
_c_, Outer wall.
_n_, Nucleus, or primary chamber.
_v_, Internal cavity.]
Receptaculites is a still more complex organism. It has a sack-like
form, often attaining a large size, and the double walls are composed of
square or rhombic plates, connected with each other by hollow tubes from
which proceed canals perforating the plates (Fig. 25). This curious
structure is confined to the Siluro-Cambrian, and is so dissimilar from
modern forms that its affinities have been subject to grave doubts.
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