The Chain of Life in Geological Time: A Sketch of the Origin and Succession of Animals and PlantsDawson, John William, Sir
Science
The Chain of Life in Geological Time: A Sketch of the Origin and Succession of Animals and Plants
Dawson, John William, Sir
Paleobotany; Paleontology
The Foraminifera of the Palæozoic we have noticed in the last chapter;
but we now find a new type of Protozoan—that of the Sponge. Sponges as
they exist at present may be defined to be composite animals, made up of
a great number of one-celled or gelatinous zoids, provided with
vibrating threads or cilia, and so arranged that currents of water are
driven through passages or canals in the mass, by the action of the
cilia, bringing food and aerated water for respiration. To support these
soft sarcodic sponge-masses, they secrete fibres of horny matter and
needles (spiculæ) of flint or of limestone, forming complicated fibrous
and spicular skeletons, often of great beauty. They abound in all seas,
and some species are found in fresh waters.
[Illustration: FIG. 29.—Portion of skeleton of Hexactinellid Sponge
(_Cœloptychium_). Magnified. After Zittel.]
With the exception of a very few species destitute of skeleton, and
which we cannot expect to find in a fossil state, the sponges may be
roughly divided into three groups: 1, those with corneous or horny
skeleton, like our common washing sponges; 2, those with skeletons
composed of silicious needles of various forms and arrangement; 3, those
with calcareous spicules. Of these, the second or silicious group has
precedence in point of time, beginning in the Early Cambrian, and
continuing to the present. Two of its subdivisions are especially
interesting in their range. The first is that of the Lattice-sponges
(_Hexactinellidæ_), in which the spicules have six rays placed at right
angles, and are attached to each other by their points, so as to form a
very regular network (Fig. 29). The second is that of the Stone-sponges
(_Lithistidæ_), in which the spicules are four-rayed or irregular, and
are united by the branching root-like ends of the rays. The most
beautiful of all sponges, the Venus Flower-basket (_Euplectella_), is a
modern Hexactinellid, and the wonderful weaving of its spicules is as
marvellous a triumph of constructive skill as its general form is
graceful. The Lithistids are less beautiful, but are the densest and
most compact of sponges, and are represented by several species in the
modern seas. Both of these types go back to the Early Cambrian, and have
continued side by side to the present day, as representatives of two
distinct geometrical methods for the construction of a spicular
skeleton.
[Illustration: FIG. 30.—_Protospongia fenestrata_ (Salter). Menevian
group.
_a_, Fragment showing the spicules partially displaced. _b_, Portion
enlarged.]
[Illustration: FIG. 31.—_Astylospongia præmorsa_ (Roemer). Niagara
group.—After Hall.
_a_, Spicules magnified.]
[Illustration: FIG. 32.—Spicules of Lithistid sponge (_Trichospongia_
of Billings). From the Cambrian of Labrador.]
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