The Chain of Life in Geological Time: A Sketch of the Origin and Succession of Animals and PlantsDawson, John William, Sir
Science
The Chain of Life in Geological Time: A Sketch of the Origin and Succession of Animals and Plants
Dawson, John William, Sir
Paleobotany; Paleontology
The true Stony Corals (_Anthozoa_) are as yet unknown in the Cambrian.
They entered on the stage in immense abundance in the Siluro-Cambrian,
where considerable limestones are largely composed of their remains,
mixed, however, and sometimes overpowered with those of Bryozoa and
Hydroids. An ordinary coral, such as those of which coral reefs are
built—the red coral, used for ornament is not quite similar—is the
skeleton of an animal constructed on the plan of a sea anemone; with a
central stomach surrounded by radiating chambers, and having above a
crown of tentacles. The stony coral surrounds and protects the soft body
of the animal, and may either be a single cell, for one animal, or an
aggregation of such cells, constituting a rounded or branching mass.
The modern star coral, represented in Fig. 42, is an instance of the
latter condition. It shows nineteen or twenty animals, each with a
central mouth and fringe of short tentacles, aggregated together, and
two of them showing the spontaneous division by which the number of
animals in the mass is progressively increased. The living coral shows
only the soft animals and the animal matter connecting them; but if dead
there would be a white stony mass with a star-like cell or depression
corresponding to each animal.
[Illustration: FIG. 42.—Living Anthozoan Coral (_Astræa_).]
In their general plan, the oldest Corals were precisely of this
character, but they presented some differences in detail, which have
caused them to be divided into two groups, which are eminently
characteristic of the Palæozoic age—the tabulate or floored corals, and
the rugose or wrinkled corals. In the former (Fig. 43) the cells are
usually small and thin-walled, often hexagonal, like a honeycomb, and
are floored across at intervals with tabulæ or horizontal plates. A few
modern corals present a similar arrangement,[14] but this kind of
structure was far more prevalent in the Palæozoic. In the second type
the animals are usually larger and often solitary, the cell has strongly
marked radiating plates, while the horizontal floors are absent or
subordinate, and there is usually a thick external rind or outer coat
(Figs. 44, 45). In general plan, these rugose corals closely resemble
those of our modern reefs; but they differ in their details of
structure, and only a very few modern forms from the deep sea are
regarded as actual modern representatives.[15] One curious point of
difference is that their radiating laminæ begin with four, and increase
by multiples of that number, while in modern corals the numbers are six
and multiples of six; a change of mathematical relation not easily
accounted for, and which assimilates them to Hydroids on the one hand,
and to a higher group, the Alcyonids, on the other, both of which prefer
four and eight to six, or have had these numbers chosen for them. In the
Mesozoic period the tabulate and rugose corals were replaced by others,
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