The Chemistry of Plant LifeThatcher, Roscoe Wilfred
Science
The Chemistry of Plant Life
Thatcher, Roscoe Wilfred
Botanical chemistry
While the proteins are finally stored up largely in the seeds, or other
storage organs, they are actively at work during the growing period in the
cells of the foliage parts of the plant. Hence, the popular statement that
"nitrogen makes foliage" is a fairly accurate expression of its rôle.
Inordinate production of straw in cereal crops and of leaves in root crops
often results from liberal supplies of available nitrogen in the soil early
in the growing season. If the crops develop to normal maturity, this
excessive foliage growth has no harmful results, as the surplus material
which has been elaborated is properly translocated into the desired storage
organs; but, unfortunately, the retarding effect of the surplus nitrogen
supply upon the date of maturing of the crop is often associated with
premature ripening of the plants from other causes, with the consequence
that too large a proportion of the valuable food material is left in the
refuse foliage material of the crop. Crops which are grown solely for their
leaves, such as hay crops, lettuce, cabbage, etc., profit greatly by
abundant supplies of available nitrogen; although when foliage growth is
stimulated in this way the tissue is likely to be thin-walled and soft
rather than firm and solid.
=Phosphorus= is likewise an extremely important element in plant nutrition.
But phosphorus starvation produces no such striking visible effects upon
the growth of the plant as does lack of nitrogen. Abundance of available
phosphorus early in the plant's life greatly stimulates root growth, and
later on it undoubtedly hastens the ripening process; hence, this element
seems to act as the exact antithesis of nitrogen.
The rôle of phosphorus, or of phosphates, in the physiological processes of
the cell seems to be difficult to discover. The element itself is a
constituent of some protein complexes and of the lecithin-like bodies (see
page 141) which are supposed by some investigators to play an important
part in determining the rate of chemical changes which take place in the
cell and the movement of materials into and out of it. It is an essential
constituent of the nucleus, and a meager supply of phosphorus retards, or
inhibits, mitotic cell-division. Photosynthesis of sugars and the
condensing of these into starch or cellulose takes place in plants in the
absence of available phosphorus; but the change of these insoluble
carbohydrates back again into soluble and available sugar foods does not.
Phosphorus is taken from the soil by plants in the form of phosphates. Much
study has been given to the problem of the proper supply of available soil
phosphates for economic crop production. Any discussion of soil fertility
and fertilization which did not devote large attention to the conditions
under which phosphates become available as plant food would be wholly
inadequate; but such a discussion would be out of place here.
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