The Chemistry of Plant LifeThatcher, Roscoe Wilfred
Science
The Chemistry of Plant Life
Thatcher, Roscoe Wilfred
Botanical chemistry
Much study has been given during recent years to the question of the
supposed poisonous, or toxic, effects upon plants of various soil
constituents. There seems to be no doubt that certain _organic_ compounds
which are injurious to plant life are often present in the soil, either as
the normal excretions of plant roots or as products of the decomposition of
preceding plant growths. A consideration of these supposedly toxic organic
substances would be out of place in this discussion of mineral soil
nutrients. But there seems to be no doubt that there may also be mineral
substances in the soil which may sometimes exert deleterious influences
upon plant growth. In fact, most metallic salts, except those of the few
metals which are required for plant nutrition, appear to be toxic to
plants. The exact nature of the physiological effects which are produced by
these mineral toxins is not clearly understood; indeed, it is probably
different in the case of different metals. Further, it is certain that both
the stimulating and the toxic effect of metallic compounds upon low forms
of plants is quite different from the effects of the same substances upon
the more complex tissues of higher plants, a fact which is utilized to
advantage in the application of fungicides for the control of parasitic
growths on common farm crops.
Among the elements whose physiological effects upon higher plants, such as
the cereal crops, etc., when their soluble compounds are present in the
soil, have been carefully studied, there are three fairly distinct types of
injurious mineral elements. The first of these, represented by copper,
zinc, and arsenic, apparently exert their toxic effect regardless of the
proportion in which they are present in the nutrient solution which is
presented to the plant; although the degree of injury varies with the
amount of injurious substance present, of course. The second type, of which
boron and manganese are representatives, apparently exerts a definite
stimulating effect upon plants when supplied to them in concentrations
below certain clearly defined limits; but are toxic in concentrations above
these. The third includes many soluble salts of magnesium, sodium,
potassium, etc., which while either innocuous or else definite sources of
essential plant foods when in lower concentrations, become highly toxic, or
corrosive, when present in the soil solution in concentrations above the
limits of "toleration" of individual plants for these soluble salts. The
tolerance shown by the different species of plants toward these soluble
salts (the so-called "alkali" in soils) varies widely; indeed, there seems
to be considerable variation in the resistance of different individual
plants of the same species to injury from this cause.
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