The Coal Measures Amphibia of North AmericaMoodie, Roy Lee
Science
The Coal Measures Amphibia of North America
Moodie, Roy Lee
Amphibians, Fossil -- North America; Paleontology -- Carboniferous; Paleontology -- North America
There are four elements which take part in the formation of the posterior
border of the skull. These are the postparietal, the tabulare, the
squamosal, and the supratemporal. It is very unusual for the supratemporal
to reach the posterior edge of the cranium. The pref rental lies anterior
to the orbit, of which it forms the anterior border. The lacrimal has not
been detected, although Jaekel (347) has indicated it in his drawings
of the skull of the type species. The maxilla is elongate and forms the
lateral border of the skull. No teeth have been detected, although they
were doubtless the same as Jaekel has figured in _D. punctolineatus_.
The jugal is an elongate element joining the maxilla posteriorly. Jaekel
included this element in his "perisquamosal," but the sutures are clearly
evident in the present specimen and there is no evidence of a structure at
all similar to a "perisquamosal." The postorbital is fully as large as the
jugal which it joins, forming a part of the posterior border of the orbit
and ending posteriorly in a point which is inclosed by the tabulare and
the squamosal. The postfrontal with the foregoing element forms the entire
posterior border of the orbit and it likewise ends in a point inclosed
by the parietal and the postorbital. The quadratojugal has much the same
shape and relations as in _D. punctolineatus_, although it is located
further back. The squamosal is also elongate, as are most of the posterior
cranial elements, and it also has an acumination which is directed forward
and is inclosed by the postorbital and jugal. The anterior suture of this
element is rather indistinct, but it is, I believe, as represented (fig.
24). The element is elongate and is prolonged posteriorly to form the
horn, which ends in a blunt point and is not sharp, as in the type species.
Jaekel (347) regards the species _Diceratosaurus punctolineatus_ Cope
as being unparalleled among known vertebrates in the possession of a
"perisquamosal" element. In closely allied species the "perisquamosal" is
easily separated into its component elements, and the morphology of the
present skull would throw considerable doubt on Jaekel's interpretation
of the skull of the type species. Another specimen, described below as
another species of this genus, shows no evidence of this fusion. So far as
I can learn, there have been no cases of true fusion of cranial elements
correctly reported, unless it be that which possibly exists between the
two frontals in the skull of _Diplocaulus_. It was on the basis of such
fusions that Maggi (397) proposed to derive the interparietals of the
primates from the tabulare of the stegocephalians.
The posterior outline of the skull in the present specimen is not well
preserved and the outline as given may be slightly inaccurate. The
indentation figured by Jaekel in the posterior border of the skull of the
type form is not present in the species under discussion.
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