The Coal Measures Amphibia of North AmericaMoodie, Roy Lee
Science
The Coal Measures Amphibia of North America
Moodie, Roy Lee
Amphibians, Fossil -- North America; Paleontology -- Carboniferous; Paleontology -- North America
The premaxillaries, which show some obscure traces of a symphysial suture
at the median line, anterior to the nasal or naso-palatine vacuities,
extend outwards, on each side, for an extent of 2.5 lines and there
join the maxillaries. Traces of round alveoli for teeth, some of which
are 2 lines in diameter, are visible on the alveolar border of the
premaxillaries. The alveolar border is continued by the maxillary bone for
an extent of 4.5 inches beyond the premaxillary border, and this border
shows still more distinct traces of alveoli, of a circular form, about a
line in diameter and rather close set in a single series. The fore part
of the orbit is very unequivocally displayed, the smooth inner or under
surface of the bone forming that part being entire; and this shows the
fore part of the orbit to be formed, partly by the maxillary, partly
by the lacrimal or prefrontal bone in close sutural union therewith,
a structure which does not exist in any recent or fossil fish with a
dentigerous superior maxillary bone. Where the substance of the bone has
been detached so far as to expose the external layer in contact with
the coal, as, _e.g._, on the frontal and part of the prefrontal bones,
the external surface of those bones is shown to have been impressed by
subhemispherical or elliptical pits, from 1 line to 1.5 lines in diameter,
and with intervals of half of that extent. This coarsely pitted character
agrees with that presented by the other surface of the similarly broad and
flat cranium of the labyrinthodonts.
From the characters above specified, therefore, I conclude that this
fossil is the fore part of the skull of an extinct family of the
labyrinthodonts. It agrees with them in the number, size, and disposition
of teeth; in the proportions and mode of connection of the premaxillaries,
maxillaries, nasals, prefrontals, and frontals, and in the resultant
peculiarly broad and depressed character of the skull. The traces of
the nostrils are less definite and satisfactory than the remains of the
orbits, but the latter appear to be decisive against the piscine nature
of the fossil. The fossil also presents the same well-marked external
sculpturing as in the labyrinthodonts; and among the genera that have
been established in that family, the form of the end of the muzzle, or
upper jaw, in the Pictou coal specimen best accords with that in the
_Capitosaurus_ and _Metopias_ of von Meyer and Burmeister (80).
Measurements of the Skull of Baphetes planiceps Owen.
(Type in the British Museum of Natural History, London.)
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