3. STARTING OF BUDS.--Deciduous trees are entirely destitute of leaves
during the winter. The buds must start to grow in the spring without
leaves, and therefore without drawing force from the sun. Hence,
also, food in the organic form must be, and is, laid up from the
previous year in the body of the tree. A portion of this is consumed
with the formation of CO₂ and H₂O, in order to create force for
the development of the buds. So soon as by this means the leaves are
formed, the plant begins to draw force from the sun, and food from the
mineral kingdom.
4. PALE PLANTS.--Fungi and etiolated plants have no chlorophyl,
therefore cannot draw their force from the sun, nor make organic
matters from inorganic. Hence these also must feed on organic matter;
not, indeed, on starch, dextrine, and protoplasm, but on decaying
organic matter. In these plants the organic matter is taken up in some
form intermediate between the planes No. 3 and No. 2. The matter thus
taken up is, a portion of it, consumed with the formation of CO₂ and
H₂O, in order to create force necessary to organize the remainder.
To illustrate: Matter falling from some intermediate point between No.
2 and No. 3 to No. 2, produces force sufficient to raise matter from
the same intermediate point to No. 3; a portion runs to waste downward,
and creates force to push the remainder upward.
5. GROWTH OF GREEN PLANTS AT NIGHT.--It is well known that almost all
plants grow at night as well as in the day. It is also known that
plants at night exhale CO₂. These two facts have not, however, as
far as I know, been connected with one another, and with the principle
of conservation of force. It is usually supposed that in the night
the decomposition of CO₂ and exhalation of oxygen are checked by
withdrawal of sun light, and some of the CO₂ in the ascending sap is
exhaled by a physical law. But this does not account for the growth. It
is evident that, in the absence of sun light, the force required for
the work of tissue-building can be derived only from the decomposition
and combustion of organic matter. There are two views as to the source
of this organic matter, either or both of which may be correct: First.
There seems to be no doubt that most plants, especially those grown in
soils rich in _humus_, take up a portion of their food in the form of
semi-organic matter, or soluble _humus_. The combustion of a portion of
this in every part of the plant, by means of oxygen also absorbed by
the roots, and the formation of CO₂, undoubtedly creates a supply of
force night and day, independently of sunlight. The force thus produced
by the combustion of a portion might be used to raise the remainder
into starch, dextrine, etc., or might be used in tissue-building.
During the day, the CO₂ thus produced would be again decomposed in
the leaves by sunlight, and thus create an additional supply of force.
During the night, the CO₂ would be exhaled.[14]
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