The Swiss naturalist, accordingly, in adopting a new principle of
classification, so essential in the case of the fossils of the old red
sandstone, has made the scales and external organs the groundwork of his
system. The classification of Cuvier and the older naturalists proceeds
mainly upon the character and disposition of the fins. Hence the order
of the Acanthopterygii, or thorny-finned; and the Malacopterygii, or the
soft finned order.—The classification of M. Agassiz, proceeding upon the
characters of the scales and plates, has given rise to the following
orders, namely, the Placoid, or broad-plated scale; the Ganoid, or the
shining-scale; the Ctenoid, or comb-shaped scale; and the Cycloid, or
marginated scale. Upon the simple basis of these four orders, he has
constructed his system and composed his “Poissons Fossiles,” the standard
authority in fossil ichthyology, and elaborate monument of his learning
and genius.
The relations, as well as distinguishing peculiarities, of the fishes
of the old red, are thus described by Agassiz:—“Of the Placoidian
order,” he says, “the genera ctenacanthus, onchus, ctenoptychius,
and ptychacanthus, are provided with spinous rays to the dorsal fins,
resembling the gigantic ichthyodorulites of the carboniferous and
jurassic formations, but differing in their less considerable size; they
are distinguished among themselves by the forms and ornaments of their
rays. In the order of ganoid fishes, the genera acanthodes, diplacanthus,
cheiracanthus, and cheirolepis present themselves at first sight as a
separate group; for although covered, like the others, with enameled
scales, these are so small, that they impart to the skin the appearance
of shagreen. The manner in which the fins are sustained by spinous rays,
or the absence of these rays, and the position of the fins themselves,
have served as characters in the establishing of these genera. The genera
pterichthys, coccosteus, and cephalaspis, form a second group exceedingly
curious: the considerable development of the head, its size, large plates
which cover it, and which likewise extend over the greater portion of
the trunk, and the movable appendages in the form of a wing, placed on
the side of the head, give to them the most remarkable appearance. It is
these peculiarities, indeed, which caused the class to which these genera
belong for a long time to be misunderstood. The large bony and granulated
plates of coccosteus, led to their being considered as belonging to
trionyx: and it will be sufficient excuse for this error to call to
recollection, that the greatest anatomist of our age had sanctioned this
approximation. The form of the disc of the head of the cephalaspides,
which has the appearance of a large crescent, and their more numerous,
but very elevated scales, resembling the transverse articulations of the
body, explain how it was possible to see in these fishes the trilobites
of a particular genus. Lastly, the winged appendages of the sides of
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