In the interradii the marginal lamella undergoes modifications due to the
fact that the bases of the pedalia are situated a little upwards from
the exact margin, and that the lamella follows the outline of the bases.
Fig. 1 shows one of the interradial corners of the bell margin looked at
directly from the surface, so that the curved outline of the junction of
the base of the pedalium with the exumbrella is seen. The trace made by
the lamella where it meets the surface ectoderm follows this outline. The
lamella is also shown in the vertical section through the interradius
(Fig. 5 or 20, _vlm_), where it is seen running from the connecting
canals (_cc_), which joins the two adjacent marginal pockets, upwards and
outwards to meet the surface ectoderm. Its course from canal to surface
is not in a direct line, but curved with the concavity upwards. Hence, in
cross-sections at certain levels through the interradial corner it is met
more than once and gives rise to appearances that seem at first sight too
complicated for it to be just the same structure as the simple marginal
lamella described above. That it is the same, and that the complication
is only due to the insertion of the pedalia above the margin, can be
determined by following through a series of cross-sections, the essential
ones of which, as I hope, are given in Figs. 40-43. The levels of these
are shown on Fig. 5 by the letters _w_, _x_, _y_ and _z_, respectively.
Fig. 40 shows the lamella cut but once, just below its highest part. The
section is above the level of the connecting canal and hence still shows
the vertical interradial lamella _ivl_. Fig. 41, at the next lower level
(_x_), shows the same portion of the lamella intersected a little nearer
the interior, while the junction with the endoderm of the connecting
canal is shown still further inside. Fig. 42 is at level _y_, just
through the bend of the loop, so that in part of its course the lamella
is cut almost horizontally, _i. e._ in its own plane. Fig. 43 finally
shows the lamella as it appears below the level of the connecting canal,
cut twice, each portion joining endoderm of marginal pocket with ectoderm
of surface. It thus bears exactly the same relations that it had when we
first met it in Fig. 38 (_vlm_), except that here in Fig. 43 one finds
that a cross-section cuts it at right angles instead of a vertical as in
Fig. 38, as a result of its being pushed upwards from its former position
on the margin by the insertion of the pedalium above the margin.
Public-domain text, read in full here on John Shaqi.
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