(b) Yet the fibres that Schewiakoff observed and associated with special
visual cells occur beyond question. Fig. 64 is a drawing of the first
cut through the vitreous body of Charybdea, and in among the sections of
the pigment streaks are seen sections of processes lying within clear
spaces exactly as Schewiakoff figures his visual fibres (’89, Taf. II,
Fig. 18). That the fibres occur is indisputable, but as to the cells to
which they belong I can say nothing except that from such evidence as
I have given in the preceding paragraph I conclude that they come from
pigmented retinal cells of not very different type within the retina from
the others, if different at all.
(c) On the third point, that the pigment streaks in the vitreous body
belong to underlying cells and are continued distally into fibrous
processes like the visual fibres of Schewiakoff, the evidence is
decisive. Fig. 58 has already shown it, and if this were not enough, a
case of unusual stoutness of the fibres drawn in Fig. 67 is conclusive.
The preparation from which the section is taken was one preserved with
corrosive-acetic, and I have drawn the outlines with the camera in
order to avoid exaggeration of the fibres as far as possible, and also
to show the shrinkage of the vitreous body (_vb_). It is the shrinkage
of the vitreous body that makes it so difficult to determine the exact
relation of structures seen in the vitreous body to the retina. The
fibrous processes run through the vitreous body to the “capsule” of the
lens (_cp_) (see also Fig. 72), a layer of homogeneous substance much
resembling that of the vitreous body, which is classed as a part of
the vitreous body, but usually in the shrinking adheres to the lens.
The capsule is therefore regarded by Schewiakoff as a secretion of the
lens cells. Some fibres were found by him to have the appearance of
branching upon reaching the surface of the capsule, others of passing
through it and of seemingly ending among the cells of the lens. The
same appearances were given in my sections. It is altogether impossible
in the distal portion of the vitreous body to distinguish between the
fibres of Schewiakoff and those that come from the long pigment cells.
(Figs. 64-66 represent the appearance of the vitreous body at successive
levels, and are from the same series of sections as Figs. 59-62 and 72.)
In Fig. 64 the sections of the processes that Schewiakoff calls visual
are easily distinguished from the sections of the long pigment cells.
In Fig. 65, which is two or three sections nearer the lens, the pigment
cells are shown by their cross-sections to be tapering down, and in Fig.
66, nearer still to the lens, the two kinds of processes are no longer
to be distinguished from each other. In a few cases I have found pigment
in a fibre which but for this would be called one of the visual fibres
of Schewiakoff. Such considerations as these, the similar appearance in
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