The Cubomedusæ are of more than passing interest among jelly-fish,
both because of their comparative rarity and because of the high
degree of development attained by their nervous system. One fact alone
suffices to attract at once the attention of the student of comparative
morphology--that here among the lowly-organized Cœlenterates we find an
animal with eyes composed of a cellular lens contained in a pigmented
retinal cup, in its essentials analogous to the vertebrate structure.
Perhaps this and other facts about the Cubomedusæ would be more generally
known, had they not been to a certain extent hidden away in Claus’s
paper on Charybdea marsupialis (’78), which, while a record of careful
and accurate work, is in many respects written and illustrated so
obscurely that it is very doubtful whether one could arrive at a clear
understanding of its meaning who was not pretty well acquainted with
Charybdea beforehand.
Before Claus’s paper was received at this laboratory, H. V. Wilson went
over essentially the same ground upon a species of Chiropsalmus taken
at Beaufort, N. C. When the article on Charybdea marsupialis appeared,
however, the results were so similar that Wilson did not complete for
publication the careful notes and drawings he had made.
Haeckel’s treatment of the Cubomedusæ in his “System” (’79) in the
Challenger Report (’81) is much more lucid than Claus’s; but the extended
scope of his work and the imperfect preservation of his material
prevented a detailed investigation, and for a more complete and readily
intelligible account of the structure of the Cubomedusæ a larger number
of figures is desirable.
In the foregoing facts lies whatever excuse is necessary for repeating
in the present paper much that has already seen print in one form or
another.
PART I: SYSTEMATIC.
It seems advisable first of all to establish the systematic position
of the two newly found species, Charybdea Xaymacana and Tripedalia
cystophora. Haeckel’s classification, as given in his “System der
Medusen,” is an excellent one and will be followed in this case. One of
the new species, however, will not classify under either of Haeckel’s
two families, so that for it a new family has been formed and named the
Tripedalidæ. In showing the systematic position of the two new forms, an
outline of Haeckel’s classification will be given, so far as it concerns
our species, together with the additions that have been made necessary.
CUBOMEDUSÆ (Haeckel, 1877).
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