The Deseado Formation of PatagoniaLoomis, Frederic Brewster
Science
The Deseado Formation of Patagonia
Loomis, Frederic Brewster
Paleontology -- Patagonia (Argentina and Chile)
[Illustration: Fig. 21. (Half tone.) Restoration of Protheosodon
coniferus to show the relatively short limbs, and heavy build as
compared with Theosodon—⅕ natural size.]
RESTORATION
In order to get a comparison of what is known of this form with
_Theosodon_ I have outlined a restoration of the animal as a whole,
realizing that some essential parts are lacking, but the general
proportions can hardly vary greatly from those given. It appears,
first, that this form has an unusually short back. Though the limbs and
lower jaw are ⅗ the length of those of _Theosodon garrettorum_, the
vertebrae are ½ as long. I have assumed that the number of vertebrae
would prove to be the same as in _Theosodon_. While the limb bones
are ⅗ as long as in the _Theosodon_, they are relatively half again
as heavy and with the processes much more developed. The greatest
difference is found in the tarsus which is only ⅓ as long as that of
_Theosodon_, though relatively as heavy, and the foot was carried in
a nearly plantigrade position the heel raised but a little from the
ground, though the anticular ends of the metatarsals and the phalanges
indicate that there was a considerable freedom of movement of the
various elements. The form seems to be fairly close to the ancestral
types such as _Lambdaconus_ of the Casamayor, the limbs of which,
however, are entirely unknown, but I should expect that when found
these earlier forms would prove to be approximately plantigrade.
=Coniopternium= Ameghino
Coniopternium Amegh., 1895 Bol. Inst. Geog. Argen., t. 15, p. 632.
Coniopternium Amegh., 1897, Bol. Inst. Geog. Argen., t. 18, p. 453.
The genus is based on a calcaneum and astragulus of the macrauchenid
type, but of unusually large size. The real generic characters are
not evident in the description, but the presence of these bones,
and of three cervical vertebrae, which we also found, indicating a
macrauchenid of about the same size, are evidence that a form larger
than the Santa Cruz representatives will turn up in the Deseado beds,
for which this name may be reserved. The material is described under
the specific name _C. andinum_.
=Adianthidae= Ameghino
This family is based primarily on the genus Adianthus of the Santa
Cruz to contain some macrauchenid-like forms which, however, are of
much smaller size, and differentiated by the narrow character of the
teeth and their early tendency to hypsodonty. It seems to be a valid
series of dwarf types, which are all scarce and known only by the
most fragmentary remains. Two genera are described from the Deseado,
_Tricoelodus_, peculiar in having the posterior lobe of the lower
molars somewhat subdivided so that the tooth appears three-lobed; and
_Proadianthus_, known only by premolars which however show an unusual
development of the styles on the inner side of the teeth.
=Tricoelodus= Ameghino
Tricoelodus Amegh., 1897, Bol. Inst. Geog. Argen., t. 18, p. 454.
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