The Elements of Botany, For Beginners and For SchoolsGray, Asa
Science
The Elements of Botany, For Beginners and For Schools
Gray, Asa
Plants
[Illustration: Fig. 21. Fruit (one key) of Silver Maple, Acer
dasycarpum, of natural size, the seed-bearing portion divided to show
the seed. 22. Embryo of the seed taken out. 23. Same opened out, to show
the thick cotyledons and the little plumule or bud between them. 24.
Germination of Silver Maple, natural size; merely the base of the fruit,
containing the seed, is shown. 25. Embryo of same, taken out of the
husk; upper part of growing stem cut off, for want of room.]
24. =Examples of Embryos with thickened Cotyledons.= In the Pumpkin and
Squash (Fig. 26, 27), the cotyledons are well supplied with nourishing
matter, as their sweet taste demonstrates. Still, they are flat and not
very thick. In germination this store is promptly utilized in the
development of the caulicle to twenty or thirty times its length in the
seed, and to corresponding thickness, in the formation of a cluster of
roots at its lower end, and the early production of the incipient
plumule; also in their own growth into efficient green leaves. The case
of our common Bean (Phaseolus vulgaris, Fig. 28-30) is nearly the same,
except that the cotyledons are much more gorged; so that, although
carried up into the air and light upon the lengthening caulicle, and
there acquiring a green color, they never expand into useful leaves.
Instead of this, they nourish into rapid growth the plumule, which is
plainly visible in the seed, as a pair of incipient leaves; and these
form the first actual foliage.
25. Very similar is the germination of the Beech (Fig. 31-33), except
that the caulicle lengthens less, hardly raising the cotyledons out of
the ground. Nothing would be gained by elevating them, as they never
grow out into efficient leaves; but the joint of stem belonging to the
plumule lengthens well, carrying up its pair of real foliage-leaves.
26. It is nearly the same in the Bean of the Old World (Vicia Faba, here
called Horse Bean and Windsor Bean): the caulicle lengthens very little,
does not undertake to elevate the heavy seed, which is left below or
upon the surface of the soil, the flat but thick cotyledons remaining
in it, and supplying food for the growth of the root below and the
plumule above. In its near relative, the Pea (Fig. 34, 35), this use of
cotyledons for storage only is most completely carried out. For they are
thickened to the utmost, even into hemispheres; the caulicle does not
lengthen at all; merely sends out roots from the lower end, and develops
its strong plumule from the upper, the seed remaining unmoved
underground. That is, in technical language, the germination is
_hypogæous_.
[Illustration: Fig. 26. Embryo of Pumpkin-seed, partly opened. 27. Young
seedling of same.]
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