Anatomy, Comparative; Embryology, Human; Evolution; Human beings -- Origin
Man is like all the other higher animals, especially the apes, in
regard to these earliest and most important processes. As the human
embryo does not essentially differ, even at a much later stage of
development—when we already perceive the cerebral vesicles, the eyes,
ears, gill-arches, etc.—from the similar forms of the other higher
mammals, we may confidently assume that they agree in the earliest
embryonic processes, segmentation and the formation of germinal layers.
This has not yet, it is true, been established by observation. We have
never yet had occasion to dissect a woman immediately after
impregnation and examine the stem-cell or the segmentation-cells in her
oviduct. However, as the earliest human embryos we have examined, and
the later and more developed forms, agree with those of the rabbit,
dog, and other higher mammals, no reasonable man will doubt but that
the segmentation and formation of layers are the same in both cases.
But the special form of segmentation and layer formation which we find
in the mammal is by no means the original, simple, palingenetic form.
It has been much modified and cenogenetically altered by a very complex
adaptation to embryonic conditions. We cannot, therefore, understand it
altogether in itself. In order to do this, we have to make a
_comparative_ study of segmentation and layer-formation in the animal
world; and we have especially to seek the original, _palingenetic_ form
from which the modified _cenogenetic_ (see p. 4) form has gradually
been developed.
This original unaltered form of segmentation and layer-formation is
found to-day in only one case in the vertebrate-stem to which man
belongs—the lowest and oldest member of the stem, the wonderful
lancelet or amphioxus (cf. Chapters XVI and XVII). But we find a
precisely similar palingenetic form of embryonic development in the
case of many of the invertebrate animals, as, for instance, the
remarkable ascidia, the pond-snail (_Limnæus_), and arrow-worm
(_Sagitta_), and many of the echinoderms and cnidaria, such as the
common star-fish and sea-urchin, many of the medusæ and corals, and the
simpler sponges (_Olynthus_). We may take as an illustration the
palingenetic segmentation and germinal layer-formation in an eight-fold
insular coral, which I discovered in the Red Sea, and described as
_Monoxenia Darwinii._
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