Anatomy, Comparative; Embryology, Human; Evolution; Human beings -- Origin
The impregnated ovum of this coral (Fig. 29 A, B) first splits into two
equal cells (C). First, the nucleus of the stem-cell and its central
body divide into two halves. These recede from and repel each other,
and act as centres of attraction on the surrounding protoplasm; in
consequence of this, the protoplasm is constricted by a circular
furrow, and, in turn, divides into two halves. Each of the two
segmentation-cells thus produced splits in the same way into two equal
cells. The four segmentation-cells (grand-daughters of the stem-cell)
lie in one plane. Now, however, each of them subdivides into two equal
halves, the cleavage of the nucleus again preceding that of the
surrounding protoplasm. The eight cells which thus arise break into
sixteen, these into thirty-two, and then (each being constantly halved)
into sixty-four, 128, and so on.[16] The final result of this
repeated cleavage is the formation of a globular cluster of similar
segmentation-cells, which we call the mulberry-formation or morula. The
cells are thickly pressed together like the parts of a mulberry or
blackberry, and this gives a lumpy appearance to the surface of the
sphere (Fig. E).[17]
[16] The number of segmentation-cells thus produced increases
geometrically in the original gastrulation, or the purest palingenetic
form of cleavage. However, in different animals the number reaches a
different height, so that the morula, and also the blastula, may
consist sometimes of thirty-two, sometimes of sixty-four, and
sometimes of 128, or more, cells.
[17] The segmentation-cells which make up the morula after the close
of the palingenetic cleavage seem usually to be quite similar, and to
present no differences as to size, form, and composition. That,
however, does not prevent them from differentiating into animal and
vegetative cells, even during the cleavage.
Gastrulation of a coral. Fig. 29—Gastrulation of a coral (_Monoxenia
Darwinii_). A, B, stem-cell (cytula) or impregnated ovum. In Figure A
(immediately after impregnation) the nucleus is invisible. In Figure B
(a little later) it is quite clear. C two segmentation-cells. D four
segmentation-cells. E mulberry-formation (morula). F blastosphere
(blastula). G blastula (transverse section). H depula, or hollowed
blastula (transverse section). I gastrula (longitudinal section). K
gastrula, or cup-sphere, external appearance.)
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