Anatomy, Comparative; Embryology, Human; Evolution; Human beings -- Origin
[2] Palingenesis = new birth, or re-incarnation (_palin_ = again,
_genesis_ or _genea_ = development); hence its application to the
phenomena which are recapitulated by heredity from earlier ancestral
forms. Cenogenesis = foreign or negligible development (_kenos_ and _
genea_); hence, those phenomena which come later in the story of life
to disturb the inherited structure, by a fresh adaptation to
environment.—Translator.
This careful discrimination between the primary or palingenetic
processes and the secondary or cenogenetic is of great importance for
the purposes of the scientific history of a species, which has to draw
conclusions from the available facts of embryology, comparative
anatomy, and paleontology, as to the processes in the formation of the
species in the remote past. It is of the same importance to the student
of evolution as the careful distinction between genuine and spurious
texts in the works of an ancient writer, or the purging of the real
text from interpolations and alterations, is for the student of
philology. It is true that this distinction has not yet been fully
appreciated by many scientists. For my part, I regard it as the first
condition for forming any just idea of the evolutionary process, and I
believe that we must, in accordance with it, divide embryology into two
sections—palingenesis, or the science of recapitulated forms; and
cenogenesis, or the science of supervening structures.
To give at once a few examples from the science of man’s origin in
illustration of this important distinction, I may instance the
following processes in the embryology of man, and of all the higher
vertebrates, as _palingenetic_: the formation of the two primary
germinal layers and of the primitive gut, the undivided structure of
the dorsal nerve-tube, the appearance of a simple axial rod between the
medullary tube and the gut, the temporary formation of the gill-clefts
and arches, the primitive kidneys, and so on.[3] All these, and many
other important structures, have clearly been transmitted by a steady
heredity from the early ancestors of the mammal, and are, therefore,
direct indications of the presence of similar structures in the history
of the stem. On the other hand, this is certainly not the case with the
following embryonic forms, which we must describe as cenogenetic
processes: the formation of the yelk-sac, the allantois, the placenta,
the amnion, the serolemma, and the chorion—or, generally speaking, the
various fœtal membranes and the corresponding changes in the blood
vessels. Further instances are: the dual structure of the heart cavity,
the temporary division of the plates of the primitive vertebræ and
lateral plates, the secondary closing of the ventral and intestinal
walls, the formation of the navel, and so on. All these and many other
phenomena are certainly not traceable to similar structures in any
earlier and completely-developed ancestral form, but have arisen simply
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