Anatomy, Comparative; Embryology, Human; Evolution; Human beings -- Origin
Fig.14 The human ovum. Fig. 14—The human ovum, taken from the female
ovary, magnified. The whole ovum is a simple round cell. The chief part
of the globular mass is formed by the nuclear yelk (_deutoplasm_),
which is evenly distributed in the active protoplasm, and consists of
numbers of fine yelk-granules. In the upper part of the yelk is the
transparent round germinal vesicle, which corresponds to the _
nucleus._ This encloses a darker granule, the germinal spot, which
shows a _nucleolus._ The globular yelk is surrounded by the thick
transparent germinal membrane (_ovolemma,_ or _ zona pellucida_). This
is traversed by numbers of lines as fine as hairs, which are directed
radially towards the centre of the ovum. These are called the
pore-canals; it is through these that the moving spermatozoa penetrate
into the yelk at impregnation.
When the mature bird-ovum has left the ovary and been fertilised in the
oviduct, it covers itself with various membranes which are secreted
from the wall of the oviduct. First, the large clear albuminous layer
is deposited around the yellow yelk; afterwards, the hard external
shell, with a fine inner skin. All these gradually forming envelopes
and processes are of no importance in the formation of the embryo; they
serve merely for the protection of the original simple ovum. We
sometimes find extraordinarily large eggs with strong envelopes in the
case of other animals, such as fishes of the shark type. Here, also,
the ovum is originally of the same character as it is in the mammal; it
is a perfectly simple and naked cell. But, as in the case of the bird,
a considerable quantity of nutritive yelk is accumulated inside the
original yelk as food for the developing embryo; and various coverings
are formed round the egg. The ovum of many other animals has the same
internal and external features. They have, however, only a
physiological, not a morphological, importance; they have no direct
influence on the formation of the fœtus. They are partly consumed as
food by the embryo, and partly serve as protective envelopes. Hence we
may leave them out of consideration altogether here, and restrict
ourselves to material points—_to the substantial identity of the
original ovum in man and the rest of the animals_ (Fig. 13).
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