Anatomy, Comparative; Embryology, Human; Evolution; Human beings -- Origin
So much for the embryonic process in itself. The application to
evolution has been a long and laborious task. Briefly, it was
necessary to show that ALL the multicellular animals passed through
these three stages, so that our biogenetic law would enable us to
recognise them as reminiscences of ancestral forms. This is the work
of Chapters 1.8 and 1.9. The difficulty can be realised in this way:
As we reach the higher animals the ovum has to take up a large
quantity of yelk, on which it may feed in developing. Think of the
bird's "egg." The effect of this was to flatten the germ (the morula
and blastula) from the first, and so give, at first sight, a totally
different complexion to what it has in the lowest animals. When we
pass the reptile and bird stage, the large yelk almost disappears (the
germ now being supplied with blood by the mother), but the germ has
been permanently altered in shape, and there are now a number of new
embryonic processes (membranes, blood-vessel connections, etc.). Thus
it was no light task to trace the identity of this process of
gastrulation in all the animals. It has been done, however; and with
this introduction the reader will be able to follow the proof. The
conclusion is important. If all animals pass through the curious
gastrula stage, it must be because they all had a common ancestor of
that nature. To this conjectural ancestor (it lived before the period
of fossilisation begins) Haeckel gives the name of the Gastraea, and
in the second volume we shall see a number of living animals of this
type ("gastraeads").
The line of argument is the same in the next chapter. After laborious
and careful research (though this stage is not generally admitted in
the same sense as the previous one), a fourth common stage was
discovered, and given the name of the Coelomula. The blastula had one
layer of cells, the blastoderm (derma = skin): the gastrula two
layers, the ectoderm ("outer skin") and entoderm ("inner skin"). Now a
third layer (mesoderm = middle skin) is formed, by the growth inwards
of two pouches or folds of the skin. The pouches blend together, and
form a single cavity (the body cavity, or coelom), and its two walls
are two fresh "germinal layers." Again, the identity of the process
has to be proved in all the higher classes of animals, and when this
is done we have another ancestral stage, the Coelomaea.
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