Anatomy, Comparative; Embryology, Human; Evolution; Human beings -- Origin
To give at once a few examples from the science of man's origin in
illustration of this important distinction, I may instance the
following processes in the embryology of man, and of all the higher
vertebrates, as palingenetic: the formation of the two primary
germinal layers and of the primitive gut, the undivided structure of
the dorsal nerve-tube, the appearance of a simple axial rod between
the medullary tube and the gut, the temporary formation of the
gill-clefts and arches, the primitive kidneys, and so on.* (* All
these, and the following structures, will be fully described in later
chapters.--Translator.) All these, and many other important
structures, have clearly been transmitted by a steady heredity from
the early ancestors of the mammal, and are, therefore, direct
indications of the presence of similar structures in the history of
the stem. On the other hand, this is certainly not the case with the
following embryonic forms, which we must describe as cenogenetic
processes: the formation of the yelk-sac, the allantois, the placenta,
the amnion, the serolemma, and the chorion--or, generally speaking,
the various foetal membranes and the corresponding changes in the
blood vessels. Further instances are: the dual structure of the heart
cavity, the temporary division of the plates of the primitive
vertebrae and lateral plates, the secondary closing of the ventral and
intestinal walls, the formation of the navel, and so on. All these and
many other phenomena are certainly not traceable to similar structures
in any earlier and completely-developed ancestral form, but have
arisen simply by adaptation to the peculiar conditions of embryonic
life (within the foetal membranes). In view of these facts, we may now
give the following more precise expression to our chief law of
biogeny: The evolution of the foetus (or ontogenesis) is a condensed
and abbreviated recapitulation of the evolution of the stem (or
phylogenesis); and this recapitulation is the more complete in
proportion as the original development (or palingenesis) is preserved
by a constant heredity; on the other hand, it becomes less complete in
proportion as a varying adaptation to new conditions increases the
disturbing factors in the development (or cenogenesis).
The cenogenetic alterations or distortions of the original
palingenetic course of development take the form, as a rule, of a
gradual displacement of the phenomena, which is slowly effected by
adaptation to the changed conditions of embryonic existence during the
course of thousands of years. This displacement may take place as
regards either the position or the time of a phenomenon.
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