Opposed to the Acrania is the second division of the vertebrates, which comprises all the other members of the stem, from the fishes up to man. All these vertebrates have a head quite distinct from the trunk, with a skull (cranium) and brain; all have a centralised heart, fully-formed kidneys, etc. Hence they are called the Craniota. These Craniotes are, however, without a skull in their earlier period. As we already know from embryology, even man, like every other mammal, passes in the earlier course of his development through the important stage which we call the chordula; at this lower stage the animal has neither vertebrae nor skull nor limbs (Figures 1.83 to 1.86). And even after the formation of the primitive vertebrae has begun, the segmented foetus of the amniotes still has for a long time the simple form of a lyre-shaped disk or a sandal, without limbs or extremities. When we compare this embryonic condition, the sandal-shaped foetus, with the developed lancelet, we may say that the amphioxus is, in a certain sense, a permanent sandal-embryo, or a permanent embryonic form of the Acrania; it never rises above a low grade of development which we have long since passed. The fully-developed lancelet (Figure 2.210) is about two inches long, is colourless or of a light red tint, and has the shape of a narrow lancet-formed leaf. The body is pointed at both ends, but much compressed at the sides. There is no trace of limbs. The outer skin is very thin and delicate, naked, transparent, and composed of two different layers, a simple external stratum of cells, the epidermis, and a thin underlying cutis-layer. Along the middle line of the back runs a narrow fin-fringe which expands behind into an oval tail-fin, and is continued below in a short anus-fin. The fin-fringe is supported by a number of square elastic fin-plates.