Anatomy, Comparative; Embryology, Human; Evolution; Human beings -- Origin
During the Triassic and Jurassic periods the sub-class of the
Monotremes was represented by a number of different stem-mammals.
Numerous fossil remains of them have lately been discovered in the
Mesozoic strata of Europe, Africa, and America. To-day there are only
two surviving specimens of the group, which we place together in the
family of the duck-bills, Ornithostoma. They are confined to Australia
and the neighbouring island of Van Diemen's Land (or Tasmania); they
become scarcer every year, and will soon, like their blood-relatives,
be counted among the extinct animals. One form lives in the rivers,
and builds subterraneous dwellings on the banks; this is the
Ornithorhyncus paradoxus, with webbed feet, a thick soft fur, and
broad flat jaws, which look very much like the bill of a duck (Figures
2.269 and 2.270). The other form, the land duck-bill, or spiny
ant-eater (Echidna hystrix), is very much like the anteaters in its
habits and the peculiar construction of its thin snout and very long
tongue; it is covered with needles, and can roll itself up like a
hedgehog. A cognate form (Parechidna Bruyni) has lately been found in
New Guinea.
These modern Ornithostoma are the scattered survivors of the vast
Mesozoic group of Monotremes; hence they have the same interest in
connection with the stem history of the Mammals as the living
stem-reptiles (Hatteria) for that of the reptiles, and the isolated
Acrania (Amphioxus) for the phylogeny of the Vertebrate stem.
The Australian duck-bills are distinguished externally by a toothless
bird-like beak or snout. This absence of real bony teeth is a late
result of adaptation, as in the toothless Placentals (Edentata,
armadillos and ant-eaters). The extinct Monotremes, to which the
Promammalia belonged, must have had developed teeth, inherited from
the reptiles. Lately small rudiments of real molars have been
discovered in the young of the Ornithorhyncus, which has horny plates
in the jaws instead of real teeth.
(FIGURE 2.269. The Ornithorhyncus or Duck-mole. (Ornithorhyncus
paradoxus).
FIGURE 2.270. Skeleton of the Ornithorhyncus.)
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