Anatomy, Comparative; Embryology, Human; Evolution; Human beings -- Origin
The living Ornithostoma and the stem-forms of the Marsupials (or
Didelphia) must be regarded as two widely diverging lines from the
Promammals. This second sub-class of the Mammals is very interesting
as a perfect intermediate stage between the other two. While the
Marsupials retain a great part of the characteristics of the
Monotremes, they have also acquired some of the chief features of the
Placentals. Some features are also peculiar to the Marsupials, such as
the construction of the male and female sexual organs and the form of
the lower jaw. The Marsupials are distinguished by a peculiar
hook-like bony process that bends from the corner of the lower jaw and
points inwards. As most of the Placentals have not this process, we
can, with some probability, recognise the Marsupial from this feature
alone. Most of the mammal remains that we have from the Jurassic and
Cretaceous deposits are merely lower jaws, and most of the jaws found
in the Jurassic deposits at Stonesfield and Purbeck have the peculiar
hook-like process that characterises the lower jaw of the Marsupial.
On the strength of this paleontological fact, we may suppose that they
belonged to Marsupials. Placentals do not seem to have existed at the
middle of the Mesozoic age--not until towards its close (in the
Cretaceous period). At all events, we have no fossil remains of
indubitable Placentals from that period.
The existing Marsupials, of which the plant-eating kangaroo and the
carnivorous opossum (Figure 2.272) are the best known, differ a good
deal in structure, shape, and size, and correspond in many respects to
the various orders of Placentals. Most of them live in Australia, and
a small part of the Australian and East Malayan islands. There is now
not a single living Marsupial on the mainland of Europe, Asia, or
Africa. It was very different during the Mesozoic and even during the
Cenozoic age. The sedimentary deposits of these periods contain a
great number and variety of marsupial remains, sometimes of a colossal
size, in various parts of the earth, and even in Europe. We may infer
from this that the existing Marsupials are the remnant of an extensive
earlier group that was distributed all over the earth. It had to give
way in the struggle for life to the more powerful Placentals during
the Tertiary period. The survivors of the group were able to keep
alive in Australia and South America because the one was completely
separated from the other parts of the earth during the whole of the
Tertiary period, and the other during the greater part of it.
(FIGURE 2.271. Lower jaw of a Promammal (Dryolestes priscus), from the
Jurassic of the Felsen strata. (From Marsh.))
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