Anatomy, Comparative; Embryology, Human; Evolution; Human beings -- Origin
If we now look back on all the remarkable features we have encountered
in the structure and the embryonic development of the Amphioxus and
the Ascidia, and compare them with the features of man's embryonic
development which we have previously studied, it will be clear that I
have not exaggerated the importance of these very interesting animals.
It is evident that the Amphioxus from the vertebrate side and the
Ascidia from the invertebrate form the bridge by which we can span the
deep gulf that separates the two great divisions of the animal
kingdom. The radical agreement of the lancelet and the sea-squirt in
the first and most important stages of development shows something
more than their close anatomic affinity and their proximity in
classification; it shows also their real blood-relationship and their
common origin from one and the same stem-form. In this way, it throws
considerable light on the oldest roots of man's genealogical tree.
CHAPTER 2.18. DURATION OF THE HISTORY OF OUR STEM.
Our comparative investigation of the anatomy and ontogeny of the
Amphioxus and Ascidia has given us invaluable assistance. We have, in
the first place, bridged the wide gulf that has existed up to the
present between the Vertebrates and Invertebrates; and, in the second
place, we have discovered in the embryology of the Amphioxus a number
of ancient evolutionary stages that have long since disappeared from
human embryology, and have been lost, in virtue of the law of
curtailed heredity. The chief of these stages are the spherical
blastula (in its simplest primary form), and the succeeding
archigastrula, the pure, original form of the gastrula which the
Amphioxus has preserved to this day, and which we find in the same
form in a number of Invertebrates of various classes. Not less
important are the later embryonic forms of the coelomula, the
chordula, etc.
Thus the embryology of the Amphioxus and the Ascidia has so much
increased our knowledge of man's stem-history that, although our
empirical information is still very incomplete, there is now no defect
of any great consequence in it. We may now, therefore, approach our
proper task, and reconstruct the phylogeny of man in its chief lines
with the aid of this evidence of comparative anatomy and ontogeny. In
this the reader will soon see the immense importance of the direct
application of the biogenetic law. But before we enter upon the work
it will be useful to make a few general observations that are
necessary to understand the processes aright.
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