Fig. 16 gives a good picture, reduced to two-thirds, of such a species,
_Phyllodes ornata_, from Assam. The posterior wings are conspicuously
coloured in deep black and yellow; in the resting position they are
covered by the anterior wings, and these are red-brown with black
markings which precisely and clearly mimic the ribs of a leaf. The
midrib begins near the tip of the anterior wing, but breaks off
half-way across the wing at two silvery white spots, similar to those
in many of the diurnal forms, which also mimic decaying leaves. Three
pairs of side veins go off backwards and forwards with remarkable
regularity from the midrib, almost at the same angle, and parallel
to one another, and three more are indicated by vague shading. Then
the midrib begins again in the internal half of the wing, though only
represented by a broad shading. The whole suggests two torn, rotten
leaves, one partly covering the other; and the deception will certainly
be perfect when the moth rests on the ground or among decaying leaves.
That all these extremely favourable protective colorations find their
explanation in the slow and gradually cumulative effects of natural
selection cannot be disputed; it is beyond doubt that they cannot be
explained, so far as we know, in any other way.
If, however, it were possible for a species of butterfly living in
the forest and among leaves to become, through natural selection,
in any degree, and in a continually increasing degree, like a leaf,
surely many insects living in the woods, and especially in the
tropical woods, would also have followed such an advantageous path of
variation--at least, so we should be inclined to think. And this is
indeed the case; numerous insects, of different orders, if they are
as large as a leaf, have taken on the colour, form, and usually also
the markings, of a leaf. Thus green and also decaying and dead leaves
are most realistically imitated by many tropical Locustidæ. Besides
_Tropidoderus_, figured on p. 79, a _Pterochroa_ of South Brazil
affords a particularly fine illustration of this, for not only does
the ground-colour, brown or green, harmonize with that of a dead or
fresh leaf, but, at the same time, all sorts of details are marked on
the insect, which help to heighten the deceptive impression. Even the
outline of the wings is leaf-like, and leaf-veins are marked on the
wing-covers with the most beautiful distinctness, and finally there
is, especially in the light-green individuals, a spot at the wing tip
which, by means of a mixture of brown, yellow, reddish, and violet
colour-tones, mimics a decaying spot with astonishing fidelity. Here,
again, the origin of this special adaptation can be clearly recognized,
for the vaguely concentric arrangement of the colours indicates
that, in the ancestors of the species, an eye-spot had occurred on
this area, of the same kind as we still see on the posterior wing,
which is covered in the resting position. Thus we can again look
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