[Illustration: FIG. 15. _Cærois chorinæus_, from the lower Amazon, in
its resting attitude. _V_, anterior wing. _H_, posterior wing. _mr_,
midrib of the apparent leaf. _sr_, lateral veins. _st_, hint of a
leaf-stalk.]
Quite different from the leaf-marking either of _Cœnophlebia_ or
_Kallima_ is that of one of the Satyrides of the lower Amazon valley,
_Cærois chorinæus_ (Fig. 15). If one spreads this butterfly out in the
usual way it does not look in the least like a leaf, and one only sees
a number of curiously placed disconnected stripes on the under surface
of the wing. But if the wings be folded together to correspond with
the sitting position of the butterfly, there appears the figure of a
leaf, of which, however, only half is present, and whose midrib (_mr_)
runs obliquely forward from the inner angle of the posterior wing.
Here, again, it is not difficult to guess that this straight stripe has
arisen, by displacement and straightening, from a curved line inherited
from some remote ancestor, and it is these precise changes which are
the work of the adaptive processes of natural selection. The same
applies to the lateral ribs (_sr_), which are here four in number.
But even the division of the wing surface by a single dark line, such
as that which crosses the middle of the posterior wing of _Hebomoja_
(Fig. 9), an Indian butterfly, heightens not inconsiderably the
resemblance of the resting butterfly to a leaf, a resemblance which
has already been shown in the form and colour. Indeed, even the sharp
division of the wing surface into a darker inner and a lighter outer
portion, which occurs in many species of _Anæa_, gives a very vivid
impression of a leaf crossed by a midrib.
It is not without a purpose that I have lingered so long over the
leaf-butterflies. I wished to make it clear that we have by no means
to do with a few exceptional cases, but with a great number, in all
of which resemblance to a leaf has been aimed at, although it has
been attained in varying degrees, and by very diverse ways. Whoever
surveys this wealth of fact must certainly receive the impression,
that, wherever it was advantageous to the existence of the species, the
evolution of such a deceptive resemblance has also been possible. In
any case one cannot but be convinced that it is not a case of chance
resemblance, as some naturalists have recently tried to maintain.
But I have not yet quite finished my outline-survey of the facts, for I
must not omit to mention that, in the evergreen tropical forests, there
are also large nocturnal Lepidoptera, which mimic leaves, sometimes
green ones, sometimes brown, dead ones.
[Illustration: FIG. 16. _Phyllodes ornata_, from Assam. Upper surface
with leaf-like marking only on the anterior wing, which is the only
part visible when at rest; ⅔ nat. size.]
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