We thus see that the facts, wherever we scrutinize them carefully,
harmonize with the theory. Of course we can only penetrate to a
certain depth with the theory of selection, and we are still far from
having reached the fundamental causes of the phenomena. Indeed, our
understanding must in the meantime stop short before the causes of
variations and their accumulation, but up to that point the theory
gives us clearness, and discloses the causal connexion of phenomena
in the most beautiful way. Although we do not yet understand how the
southern female _Papilio turnus_ was able to produce the advantageous
black, we do see why a black variation, when it did occur, should
increase and be strengthened, until it crowded out the yellow form from
the area of the immune model, and we are able in a general way to refer
the whole complicated phenomena of mimicry to their proximate causes.
This is true also of other phenomena which have had no part in
establishing the theory, since attention was only directed to them
later, and it is true even of some which, at first sight, seem to
contradict the theory altogether. To this class belongs, for instance,
the phenomenon that immune species not unfrequently mimic each other,
as was first observed among the Heliconiid-like butterflies of South
America. In four different families, the Danaidæ, the Neotropidæ, the
Heliconiidæ, and the Acræidæ, there are species, distributed over the
same area, which resemble each other in their conspicuous colouring
and marking, and also in the peculiar shape of the wings. After what
has been said one might be inclined to regard one of these species as
the unpalatable model and the others as the palatable mimics, but they
are all unpalatable, and are not eaten by birds. The puzzle of this
apparent contradiction was solved by Fritz Müller[4], who pointed out
that the aversion to non-edible butterflies is not innate in birds, but
must be acquired. Each young bird has to learn from experience which
victim is good to eat, and which bad. If every inedible species had
its particular and distinctive colour-dress a considerable number of
individuals of each species would fall victims to the experiments of
young birds in each generation, for a butterfly which has once been
pecked at, or squeezed by the bill of a bird, is doomed to die. But if
two inedible species which resemble each other inhabit the same area
they will be regarded by the birds as one and the same, and if five or
more inedible species resemble each other all five will present the
same appearance to the bird, and it will not require to repeat on the
other four the experience of unpalatability it has gained from one.
Thus the total of five species will be no more severely decimated by
the young birds than each of them would have been if it had occurred
alone; the same number of victims of experiment, which are necessary
every year in the education of the young birds, will, when all five
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