It might indeed be supposed that extermination by these could hardly
be an imminent danger, but if we think of the cockchafer blight, or of
the destruction of whole woods by the caterpillar of the 'white nun,'
or even of the destruction of several successive plantings of young
salad plants which the snails often cause in our gardens, it cannot be
doubted that all plants would be exterminated by insects and snails
alone unless they were protected against them in some degree.
We owe our detailed knowledge of the means by which plants protect
themselves against the menace of the greedy and prolific snails to
the beautiful investigations of Stahl, Professor of Botany in the
University of Jena.
In this case, too, both chemical and mechanical means are made use of.
The minute quantity of tannic acid which is contained in the leaves of
the clover prevents many snails from eating them, as, for instance,
the garden snail (_Helix hortensis_). If the leaves be soaked so as
to wash out the tannin the snail readily accepts them as food. It is
true that the small, whitish field-slug (_Limax agrestis_) does not
object to the presence of the tannin, and eats the fresh leaves of
the clover; indeed, there is no such thing as absolute protection.
In discussing the herbivorous mammals I have already mentioned that
many trees and shrubs, mosses and ferns are effectively protected by
the large amount of tannin they contain; this protection is effective
also against snails, for all these plants are fairly free from their
attacks; and the same is true of many other tannin-containing plants,
species of saxifrage and sedum, the strawberry, many water-plants, like
the pond-weeds (_Potamogeton_), the horn-nut (_Trapa_), the mare's tail
(_Hippuris_). All these plants are only eaten by snails in case of
necessity, or in the washed-out state.
In other plants protection is gained by means of some acid, especially
oxalic acid, like the wood-sorrel (_Oxalis acetosella_), the sorrel
(_Rumex_), and the species of Begonia. When Stahl smeared slices of
carrot, which is a favourite food of snails, with a weak (one per
cent.) solution of oxalate of potassium, they were refused by the
snails, and this is not surprising when we remember that even the
external skin of the snail is very sensitive, and the mucous membrane
of the mouth is not likely to be less so.
Similarly, many plants develop ethereal oils in the hairs which cover
them, as in the herb-Robert (_Geranium robertianum_). Even the almost
omnivorous field-slug (_Limax agrestis_) does not touch this plant, and
if it be placed upon it, escapes with all dispatch from the ethereal
oil, which burns its naked skin, by covering itself with mucus and
letting itself down to the ground by a thread. The mints (_Mentha_) and
the dittany (_Dictamnus albus_) also produce such oils.
Public-domain text, read in full here on John Shaqi.
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