Among chemical means of protection must be named the pure bitter
stuffs, such as are found in the species of gentian, the milkwort
(_Polygala amara_), and in many other plants, and also the curious
'oil-bodies' of the liverworts.
But some plants also defend themselves against the attacks of snails by
mechanical means.
First there are the various kinds of bristle arrangements, which
prevent the snails from creeping up the stalks. We never find the
comfrey (_Symphytum officinale_) of our meadows eaten by snails,
for it is thickly covered over with stiff bristles, which are most
disagreeable to the snail, and the stinging-nettle (_Urtica dioica_) is
similarly protected by bristle hairs, while, as we have already seen,
its stinging-hairs secure immunity from the attacks of larger animals.
And although it is true that the majority of plants do not prevent the
snails from creeping up their stalks, yet they do not serve them in any
great degree as food, since the green parts often offer resistance to
mastication and digestion. Thus the lime encrustations which cover the
stoneworts (_Chara_) prevent snails from eating them. If the lime be
dissolved by means of acids, and the plants then offered to the snails,
they will eat them greedily. The same is true of the silicifying of
the cell-walls, so widely distributed among mosses and grasses, and
when this occurs in a high degree it forms an effective protection
even against the large herbivores. Our slightly siliceous grasses are
secure from snails, and that it is really the presence of the silicic
acid which deters them from an otherwise welcome kind of food is proved
by Stahl's experiment of growing maize in pure water, and so obtaining
plants poor in silica. These were devoured without ceremony by the
snails.
Of the many other protective peculiarities which make it difficult for
snails to eat plants I shall only recall the so-called 'Raphides,'
those microscopic crystal-like needles of oxalate of lime, pointed
at both ends, which lie close together in the tissues of many
plants. Cuckoo pint (_Arum maculatum_), the narcissi, the snowdrops
(_Leucojum_), the squill (_Scilla_), and the asparagus contain them,
and all these plants are spared by snails obviously because during
mastication they are unpleasantly affected by the raphides. Even the
voracious field-slug rejects these.
Public-domain text, read in full here on John Shaqi.
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