'Instinct goes astray,' it is often said; but in truth it does not
go astray, but is only not so highly specialized in relation to the
liberating stimulus of the action as seems to us necessary for perfect
purposiveness. But in this very imperfection there lies, as it seems to
me, another proof that we have to do with the results of a process of
selection, for it is of the very nature of these never to be perfect,
but only relatively perfect, that is to say, just as perfect as is
necessary to the maintenance of the species. At the moment at which
this grade of perfection is reached every possibility of a further
increase in the effectiveness of adjustment to the end ceases, because
it would then no longer directly further the end. Why, for instance,
should the liberating stimulus in this case be more highly specialized,
since enough of the _Sitaris_ larvæ already succeed in attaching
themselves to female bees? It is not for nothing that the beetles of
this family are so prolific; what is lacking in the perfection of the
instinct is made up for by the multitude of young larvæ. A single
female of the oil-beetle (_Melöe_) lays several hundred eggs.
In speaking of the animal as a machine, it must be added that it
is a machine which can be altered in varying degrees, which can be
regulated to work at high or low pressure, slowly or quickly, finely or
roughly. This regulating is the work of the intelligence, the limited
'thinking-power,' which must be ascribed to the higher animals in a
very considerable degree, but which in the lower animals becomes less
and less apparent, until finally it is unrecognizable. That instinctive
actions can be modified or inhibited by intelligence and will is proved
by any trained beast of prey which masters its hunger and the impulse
to snap at the piece of flesh held before it, because it knows that
if it does not control itself painful blows will be the consequence.
In a later lecture I shall return to the connexion between will and
instinct; all that concerns us here is to regard instincts as the
outcome of the processes of selection, and as an indirect proof of the
reality of these.
From what I have already said at least so much must be clear, that
nothing, in principle, stands in the way of referring instincts to
selection, since their very essence is their adaptation to an end, and
such purposive changes are precisely those that are preserved in the
struggle for existence. It might, however, be supposed that in all this
the principle of use and disuse also had a share, and that without it
no changes in instincts could have come about.
There are, however, numerous instincts in considering which this can be
entirely excluded.
Public-domain text, read in full here on John Shaqi.
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