[Illustration: FIG. 42. Alpine Lousewort (_Pedicularis asplenifolia_).
_A_, flower seen from the left side, enlarged three times; the arrows
show the path by which the humble-bee enters. _B_, the same flower,
seen from the left, after removal of the calyx, the lower lip and the
left half of the upper lip. _C_, ovary (_ov_), nectary (_n_), and base
of style. _D_, tip of style, bearing the stigma. _E_, two anthers
turned towards one another. _o_, upper lip. _u_, lower lip. _gr_,
style. _st_, anthers. _kr_, corolla-tube. _k_, calyx.]
There are not a few of such flowers adapted to a very restricted
circle of visitors, and in all of them we find contrivances which
close the entrance to all except what we may call the welcome insects;
sometimes there are cushions of bristles which prevent little insects
from creeping up from below, or it is the oblique position of the
flower which prevents their getting in from the stem; sometimes it is
the length and narrowness of the corolla-tube, or the deep and hidden
situation of the nectar, which only allows intelligent insects to find
the treasure.
Very remarkable are those flowers which are adapted to the visits of
flies, for they correspond in several respects to the peculiarities
of these insects. In the first place, flies are fond of decaying
substances and the odours given off by these, and so the flowers which
depend for their cross-fertilization on flies have taken on the dull
and ugly colours of decay, and give out a disagreeable smell. But flies
are also shy and restless, turning now hither, now thither, and cannot
be reckoned among the 'constant' insect visitors, that is to say, they
do not persistently visit the same species; it is, therefore, evident
that they might easily carry away the pollen without any useful result
ensuing. Moreover, their intelligence is of a low order, and they do
not seek nectar with the perseverance shown by bees and humble-bees. It
is not surprising, therefore, to find that many of the flowers adapted
for the visits of flies are so constructed that they detain their
visitors until they have done their duty, that is to say, until they
have effected, or at least begun, the process of cross-pollination.
[Illustration: FIG. 43. Flower of Birthwort (_Aristolochia clematitis_)
cut in half. _A_, before pollination by small flies. _b_, the bristles.
_B_, after pollination. _P_, pollen mass. _N_, stigma, _b_, the
bristles. _b´_, their remains. After H. Müller.]
[Illustration: FIG. 44. Alpine Butterwort (_Pinguicula alpina_). _A_,
section through the flower. _K_, calyx. _bh_, bristly prominences.
_sp_, spur. _st_, stamen. _n_, stigma. _B_, stigma and stamen more
magnified. After H. Müller.]
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