This can be clearly demonstrated in the case of flowers. That the wild
pansy (_Viola tricolor_), which lives in the plains and on mountains of
moderate elevation, is fertilized by bees, and the nearly allied _Viola
calcarata_ of the High Alps by Lepidoptera, is readily intelligible,
since bees are very abundant in the lower region, and make the
fertilization of the species a certainty, while this is not so in the
High Alps. There the Lepidoptera are greatly in the majority, as every
one knows who has traversed the flower-decked meads of the High Alps in
July, and has seen the hundreds and thousands of butterflies and moths
which fly from flower to flower. Thus the viola of the High Alps has
become a 'butterfly-flower' by the development of its nectaries into a
long spur, accessible only to the proboscis of a moth or butterfly. The
chance which led certain individuals of the ancestral species to climb
the Alps must also have supplied the incentive to the production of the
changes adapted to the visits of the prevalent insect. The hypothesis
of a predestinating Power of Development suffers utter shipwreck in
face of facts like these.
We have, furthermore, an excellent touchstone for the reality of the
processes of selection in the _quality_ of the variations in flowers
and insects. Natural selection can only bring about those changes
which are of use to the possessors themselves; we should therefore
expect to find among flowers only such arrangements as are, directly or
indirectly, of use to them, and, conversely, among insects only such as
are useful to the insect.
Public-domain text, read in full here on John Shaqi.
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