Darwin has shown convincingly that a surprising number of characters
in animals, from worms upwards, have their roots in sexual selection,
and has pointed out the probability that this process has played an
important part in the evolution of the human race also, though, in this
case, all is not yet so clearly and certainly known as among animals.
To conclude this section, I should like once again to call attention
to the deficiency which is necessarily involved in the assumption
of any selection, sexual selection included, namely, that the first
beginning of the character which has been intensified by selection
remains obscure. Darwin attached importance to the occurrence of
ordinary individual variation, but it is open to question whether the
insignificant variations thus produced could give an adequate advantage
in the competition for the possession of the females; and, further,
whether we have not grounds for the assumption that larger variations
also occur. This question may also be asked in regard to ordinary
natural selection, although in that case we can imagine the beginnings
to be smaller, since here the advantage of a variation lies only in
the fact that it is useful, not in its being appreciated by others.
As a matter of fact, this very difficulty as to the first beginnings
of variations has been frequently urged against both hypotheses of
selection, and rightly so, inasmuch as this must be above all else
the point of attack for further investigations. But it is a mistake
to deny the whole processes of selection simply because this point is
not yet clear. Later on we shall attempt to gain some insight into
the causes of variation, and then we shall return to this question
of the beginnings of the selective processes. In the meantime let it
suffice to say that Darwin was very well aware that, in addition to the
ordinary individual variations, there were also larger deviations which
occurred discontinuously in single forms. He believed, however, that
such occurrences were very rare, and, on the whole, he was not inclined
to ascribe to them any particular importance in the transformation of
species. He rather referred the organic transformations which have
taken place in the course of the earth's history, in the main, to the
intensification of the ordinary individual variations, and I believe
that he was right in so doing, since adaptations from their very nature
cannot have been brought about by sudden chance leaps in organization,
but can only have become exactly suited to chance conditions of life
through a gradual accumulation of minute variations in the direction
of utility. Whether, however, purely sexual distinctions may not have
had their primary roots in discontinuous variations must be inquired
into later. Theoretically, there is nothing against this assumption,
when such characters are not adaptations like the lasso antennæ of
the Copepods, or the turban eyes of the Ephemerids; mere distinctive
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