markings, decorative coloration, peculiar outgrowths, and the like,
may, if they arose discontinuously, very well have formed the basis for
further sexual selection, as long as they were not prejudicial to the
existence of the species.
LECTURE XII
INTRA-SELECTION OR SELECTION AMONG TISSUES
Does the Lamarckian principle really play a part in the
transformations of species?--Darwin's position in regard to this
question--Doubts expressed by Galton and others--Neo-Lamarckians
and Neo-Darwinians--Results of exercise and practice: functional
adaptation--Wilhelm Roux, _Kampf der Theile_.
WE have devoted a whole series of lectures to studying the
Darwin-Wallace principle of Natural Selection and the range of its
operation. It seemed to us to make innumerable adaptations intelligible
up to a certain point. We now understand how the purposefulness, which
we meet with everywhere among organisms, can have arisen without
the direct interference of a Power working intentionally towards an
end--simply as the outcome and result of the survival of the fittest.
The two forms of the processes of selection, 'natural selection' in
the narrower sense, and 'sexual selection,' dominate, so to speak, all
parts and all functions of the organism, and are striving to adapt
these as well as possible to the conditions of their life. And although
the range of operation of Natural Selection is incomparably greater,
because it actually affects every part, yet we must attribute to sexual
selection also, at least among animals, a range of influence by no
means unimportant, since through it, as far as we can see at present,
not only do the secondary sexual characters in all their diversity
arise, but by the transference of these to the other sex that too is
modified, and thus the whole species may be influenced, and may indeed
be started afresh on an unlimited series of further transformations.
But although the processes of selection play such an important part in
the transformations of the forms of life, we have to inquire whether
they are the _sole_ factors in these transformations, whether the
accumulation of chance variations in the direction of utility has been
the sole factor in bringing about the evolution of the animate world,
or whether other factors have not also co-operated with it.
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