in definite sequence and arrangement, and is thus able to build up the
body of a new individual?
[Illustration: FIG. 62. _Pandorina morum_; after Pringsheim. I, A young
colony, consisting of 16 cells. II, Another colony, whose cells have
reproduced daughter-colonies; all the cells uniformly alike. III, A
young Volvox-colony; _sz_, somatic cells; _kz_, germ-cells.]
The similarity of this problem to that formulated in regard to
unicellular organisms is at once obvious, but it becomes still more
emphatic when we remember that the gulf between unicellular organisms
and the higher animals and plants is bridged over by certain transition
forms which are of the greatest interest, especially in relation to the
problems of inheritance.
Among the lower Algæ there is a family, the Volvocineæ, in which the
differentiation of the many-celled body on the principle of division of
labour has just set in; in some genera it has been actually effected,
though in the simplest way imaginable, and in others it has not yet
begun. Thus in the genus _Pandorina_ the individual consists of sixteen
green cells, united into a ball (Fig. 62, I), each one exactly like the
other, and all functioning alike. They are all united into a spherical
body, a whole, by a gelatinous matrix which they all secrete, and thus
they form a cell-colony, a cell-stock, a many-celled individual; but
each of these cells has not only all the typical parts--cell-body,
nucleus, and contractile vacuole--but each possesses a pair of flagella
or motor organs, an eye-spot, and a chlorophyll body which enables
them to assimilate nourishment from the water and the air. Each one
of these cells thus performs all the somatic functions, that is, all
that are necessary to the maintenance of the individual life. But each
also possesses the power of reproducing the whole colony from itself,
that is, it also performs the function of reproduction necessary to
the maintenance of the species. When such a colony, whose sixteen
cells are continually growing, has led for some time a free-swimming
life in the water, the cells retract their flagella, and each begins
to multiply by dividing into 2, 4, 8, finally into 16 cells of the
same kind, which remain together, forming a spherical mass enclosed in
a gelatinous secretion (Fig. 62, II). Thus there are now, instead of
sixteen cells in the mother-colony, sixteen daughter-colonies, each
with sixteen cells which soon acquire flagella and eye-spots, and are
then ready to burst forth from the dissolving jelly of the maternal
stock as independent individuals. This _Pandorina_ shows no trace of
a differentiation of its component cells to particular and different
functions, but a nearly allied genus of the same family, the genus
_Volvox_ (Fig. 62, III), consists of two kinds of cells--on the one
hand of small cells (_sz_) which occur in large numbers and compose the
wall of the hollow gelatinous mass, forming, so to speak, the skeleton
Public-domain text, read in full here on John Shaqi.
Reviews
Reviews
No reviews yet
Be the first to share your thoughts on this work.
Elsewhere in the archive
Join the Discussion
Join the discussion
Sign in to leave a comment or review.
Sign InorCreate an account