of the _Volvox_; and, on the other hand, of a much smaller number of
cells which are very much larger (_kz_). The former, the 'body' or
'somatic' cells, are green, and have a red 'eye-spot' and two flagella;
they are connected with each other by processes from their cell-bodies,
and are able, by means of the co-ordinated action of their flagella,
to propel the whole colony with a slow rotatory movement through the
water. Many of my readers are doubtless familiar with these light green
spheres, which are quite recognizable with the naked eye, and people
our marsh pools and ponds in Spring in such abundance that it is only
necessary to draw a glass of water to procure a large number of them.
The little flagellated cells just described serve not only for the
locomotion of the colony, but also for nutrition, for the secretion
of the jelly, and for the excretion of waste products; in short, they
perform all the functions necessary to the maintenance of life, but
not that of reproduction. They can, indeed, multiply by dividing when
the colony is young, like the cells of _Pandorina_, but they cannot
reproduce the whole colony but only cells like themselves, that is,
other somatic cells. In _Volvox_ the maintenance of the species, the
production of a daughter-colony, is the function of the second and
larger kind of cells, the reproductive cells, which are contained in
the interior (filled with a watery fluid) of the gelatinous sphere.
They possess no flagella (_kz_), and so take no share in the swimming
movements of the somatic cells. For the present we need not allude to
the fact that there are several kinds of these cells, and need only
state that the simplest among them, the so-called 'Parthenogonidia,'
after they have reached a considerable size, begin a process of
division which results in the formation of a daughter-colony. Usually
there are several of these large reproductive cells in a _Volvox_
colony, and as soon as these have developed into a similar number of
daughter-colonies they burst out through a rupture in the now flaccid
jelly of the maternal sphere and begin to lead an independent life. The
mother-sphere, which now consists only of somatic cells, is unable to
produce new reproductive cells; it gradually loses its spherical form,
sinks to the ground, and dies.
Public-domain text, read in full here on John Shaqi.
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